{"id":7853,"date":"2013-09-10T13:01:38","date_gmt":"2013-09-10T11:01:38","guid":{"rendered":"https:\/\/opistobranquis.info\/umbraculidaumbraculidaumbraculida"},"modified":"2026-11-05T09:10:30","modified_gmt":"2026-11-05T08:10:30","slug":"umbraculida","status":"publish","type":"page","link":"https:\/\/opistobranquis.info\/ca\/guia\/umbraculida\/","title":{"rendered":"Umbraculida"},"content":{"rendered":"<p>Els Umbraculida s\u00f3n un petit grup de mol\u00b7luscs gaster\u00f2podes heterobranquis marins caracteritzats per una conquilla externa petita i en forma de c\u00fapula, que cobreix parcialment el dors de l&#8217;animal. S\u00f3n llimacs marins de mida petita o mitjana, amb un cos tou i una morfologia particular que els diferencia d&#8217;altres heterobranquis amb conquilla. La seva anatomia combina caracter\u00edstiques pr\u00f2pies dels gaster\u00f2podes amb altres adaptacions associades a la vida bent\u00f2nica.<\/p>\n<span style=\"font-size:10pt;\" class=\"aligncenter\"><div id=\"featured-img-id\"><img data-recalc-dims=\"1\" decoding=\"async\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/12\/umbraculum-umbraculum-10.jpg?w=900&#038;ssl=1\" alt=\"Umbraculum umbraculum\" \/><\/div><center><em>Umbraculum umbraculum<\/em> per Enric Madrenas<\/center><\/span><p><\/p>\n<p>Els Umbraculida viuen principalment en ambients marins poc profunds, tot i que algunes esp\u00e8cies poden trobar-se a profunditats considerables. Es poden localitzar sobre substrats rocosos, sediments tous, entre algues o associats a altres organismes bent\u00f2nics. La seva distribuci\u00f3 \u00e9s \u00e0mplia i inclou diferents regions dels oceans tropicals, temperats i subtropicals, per\u00f2 el seu aspecte discret i els seus h\u00e0bits cr\u00edptics fan que sovint siguin dif\u00edcils de detectar.<\/p>\n<p>Una de les seves caracter\u00edstiques m\u00e9s distintives \u00e9s la conquilla, que \u00e9s petita, baixa i generalment en forma de c\u00fapula. A difer\u00e8ncia de la majoria dels gaster\u00f2podes amb conquilla, aquesta no cobreix completament el cos de l&#8217;animal, sin\u00f3 que se situa sobre la part dorsal i deixa gran part del cos exposada. El peu \u00e9s ampli i muscul\u00f3s i permet a l&#8217;animal despla\u00e7ar-se pel substrat. Al cap hi ha estructures sensorials que contribueixen a detectar l&#8217;entorn i els recursos alimentaris.<\/p>\n<p>Els Umbraculida s\u00f3n principalment herb\u00edvors o s&#8217;alimenten d&#8217;organismes s\u00e8ssils presents sobre el substrat. La seva r\u00e0dula est\u00e0 adaptada per raspar i ingerir material alimentari de les superf\u00edcies on viuen. Algunes esp\u00e8cies poden estar associades a determinats tipus d&#8217;algues o altres organismes, tot i que els coneixements sobre la seva dieta i comportament alimentari s\u00f3n encara limitats en comparaci\u00f3 amb els de grups de gaster\u00f2podes m\u00e9s estudiats.<\/p>\n<p>La morfologia dels Umbraculida reflecteix una combinaci\u00f3 de caracter\u00edstiques que els relaciona amb altres heterobranquis conquillats i amb els llimacs marins. La conquilla dorsal proporciona protecci\u00f3 sense impedir completament el moviment del cos, mentre que el peu ample permet una locomoci\u00f3 eficient sobre el substrat. Aquest pla corporal constitueix una de les formes m\u00e9s singulars d&#8217;adaptaci\u00f3 dins dels gaster\u00f2podes heterobranquis.<\/p>\n<p>Des del punt de vista evolutiu, els Umbraculida s\u00f3n especialment interessants per comprendre la diversificaci\u00f3 dels heterobranquis amb conquilla. Els estudis anat\u00f2mics i moleculars han contribu\u00eft a aclarir les seves relacions amb altres grups de gaster\u00f2podes i a comprendre millor l&#8217;evoluci\u00f3 de les seves caracter\u00edstiques particulars. La seva combinaci\u00f3 de conquilla redu\u00efda, cos tou i adaptacions bent\u00f2niques mostra la diversitat de solucions evolutives que han aparegut dins dels Heterobranchia.<\/p>\n<p>Tot i que els Umbraculida no s\u00f3n un grup habitualment observat pels bussejadors, formen part de la diversitat menys coneguda dels heterobranquis marins. La seva morfologia singular, la seva petita conquilla i la seva biologia encara poc coneguda els converteixen en un grup interessant per als estudis taxon\u00f2mics, ecol\u00f2gics i evolutius. Els mostrejos en h\u00e0bitats bent\u00f2nics poc profunds i la revisi\u00f3 detallada de substrats i organismes associats poden contribuir a millorar el coneixement sobre la seva distribuci\u00f3 i diversitat.<\/p>\n<p>La taxonomia dels Umbraculida segons WoRMS \u00e9s:<\/p>\n    <style>\n        .opk-taxonomy-loader {\n            display: flex;\n            gap: 4px;\n            align-items: center;\n            justify-content: center;\n            height: 24px;\n            margin-bottom: 1em;\n        }\n        .opk-taxonomy-loader span {\n            display: block;\n            width: 4px;\n            height: 16px;\n            background: currentColor;\n            animation: opkTaxonomyBarPulse 1.2s infinite ease-in-out;\n        }\n        .opk-taxonomy-loader span:nth-child(1) { animation-delay: 0s; }\n        .opk-taxonomy-loader span:nth-child(2) { animation-delay: 0.2s; }\n        .opk-taxonomy-loader span:nth-child(3) { animation-delay: 0.4s; }\n\n        @keyframes 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            <div class=\"autonav-image\">\n\n                            <img decoding=\"async\" width=\"650\" height=\"490\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/12\/tylodina-perversa-29.jpg?fit=650%2C490&amp;ssl=1\" class=\"autonav-thumb\" alt=\"Tylodina perversa by Enric Madrenas\" loading=\"lazy\" srcset=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/12\/tylodina-perversa-29.jpg?w=650&amp;ssl=1 650w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/12\/tylodina-perversa-29.jpg?resize=300%2C226&amp;ssl=1 300w\" sizes=\"auto, (max-width: 650px) 100vw, 650px\" data-attachment-id=\"4930\" data-permalink=\"https:\/\/opistobranquis.info\/ca\/guia\/umbraculida\/tylodina-perversa\/tylodina-perversa-29\/\" data-orig-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/12\/tylodina-perversa-29.jpg?fit=650%2C490&amp;ssl=1\" data-orig-size=\"650,490\" data-comments-opened=\"0\" data-image-title=\"Tylodina perversa by Enric Madrenas\" data-image-description=\"&lt;p&gt;Tylodina perversa by Enric Madrenas&lt;\/p&gt;\n\" data-image-caption=\"&lt;p&gt;Tylodina perversa&lt;\/p&gt;\n\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/12\/tylodina-perversa-29.jpg?fit=650%2C490&amp;ssl=1\" \/>\n                        <\/div>\n\n                    \n                    <div class=\"autonav-content\">\n\n                        <h3>\n                            Tylodina perversa                        <\/h3>\n\n                    <\/div>\n\n                <\/a>\n\n            <\/article>\n\n        \n            \n            <article class=\"autonav-card\">\n\n                <a href=\"https:\/\/opistobranquis.info\/ca\/guia\/umbraculida\/tylodina-rafinesquii\/\">\n\n                    \n                        <div class=\"autonav-image\">\n\n                            <img decoding=\"async\" width=\"810\" height=\"543\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2022\/04\/Tylodina-perversa-134-poss.-Tylodina-rafinesqui.jpg?fit=810%2C543&amp;ssl=1\" class=\"autonav-thumb\" alt=\"Tylodina rafinesquii @ by Enric Madrenas\" loading=\"lazy\" srcset=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2022\/04\/Tylodina-perversa-134-poss.-Tylodina-rafinesqui.jpg?w=810&amp;ssl=1 810w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2022\/04\/Tylodina-perversa-134-poss.-Tylodina-rafinesqui.jpg?resize=300%2C201&amp;ssl=1 300w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2022\/04\/Tylodina-perversa-134-poss.-Tylodina-rafinesqui.jpg?resize=150%2C101&amp;ssl=1 150w\" sizes=\"auto, (max-width: 810px) 100vw, 810px\" data-attachment-id=\"34022\" data-permalink=\"https:\/\/opistobranquis.info\/ca\/guia\/umbraculida\/tylodina-rafinesquii\/tylodina-perversa-134-poss-tylodina-rafinesqui\/\" data-orig-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2022\/04\/Tylodina-perversa-134-poss.-Tylodina-rafinesqui.jpg?fit=810%2C543&amp;ssl=1\" data-orig-size=\"810,543\" data-comments-opened=\"0\" data-image-title=\"Tylodina rafinesquii @ by Enric Madrenas\" data-image-description=\"&lt;p&gt;Tylodina rafinesquii @ by Enric Madrenas&lt;\/p&gt;\n\" data-image-caption=\"&lt;p&gt;Tylodina rafinesquii @&lt;\/p&gt;\n\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2022\/04\/Tylodina-perversa-134-poss.-Tylodina-rafinesqui.jpg?fit=810%2C543&amp;ssl=1\" \/>\n                        <\/div>\n\n                    \n                    <div class=\"autonav-content\">\n\n                        <h3>\n                            Tylodina rafinesquii                        <\/h3>\n\n                    <\/div>\n\n                <\/a>\n\n            <\/article>\n\n        \n            \n            <article class=\"autonav-card\">\n\n                <a href=\"https:\/\/opistobranquis.info\/ca\/guia\/umbraculida\/umbraculum-umbraculum\/\">\n\n                    \n                        <div class=\"autonav-image\">\n\n                            <img decoding=\"async\" width=\"810\" height=\"543\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/07\/Umbraculum-umbraculum-79.jpg?fit=810%2C543&amp;ssl=1\" class=\"autonav-thumb\" alt=\"Umbraculum umbraculum @ L&#039;Escala by Enric Madrenas\" loading=\"lazy\" srcset=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/07\/Umbraculum-umbraculum-79.jpg?w=810&amp;ssl=1 810w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/07\/Umbraculum-umbraculum-79.jpg?resize=300%2C201&amp;ssl=1 300w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/07\/Umbraculum-umbraculum-79.jpg?resize=150%2C101&amp;ssl=1 150w\" sizes=\"auto, (max-width: 810px) 100vw, 810px\" data-attachment-id=\"41138\" data-permalink=\"https:\/\/opistobranquis.info\/ca\/guia\/umbraculida\/umbraculum-umbraculum\/umbraculum-umbraculum-79\/\" data-orig-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/07\/Umbraculum-umbraculum-79.jpg?fit=810%2C543&amp;ssl=1\" data-orig-size=\"810,543\" data-comments-opened=\"0\" data-image-title=\"Umbraculum umbraculum @ L&amp;#8217;Escala by Enric Madrenas\" data-image-description=\"&lt;p&gt;Umbraculum umbraculum @ L&amp;#8217;Escala by Enric Madrenas&lt;\/p&gt;\n\" data-image-caption=\"&lt;p&gt;Umbraculum umbraculum @ L&amp;#8217;Escala&lt;\/p&gt;\n\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/07\/Umbraculum-umbraculum-79.jpg?fit=810%2C543&amp;ssl=1\" \/>\n                        <\/div>\n\n                    \n                    <div class=\"autonav-content\">\n\n                        <h3>\n                            Umbraculum umbraculum                        <\/h3>\n\n                    <\/div>\n\n                <\/a>\n\n            <\/article>\n\n        \n    <\/div>\n\n    \n<p><p><strong>Bibliografia<\/strong><\/p><ul><div id=\"zotpress-e23a073324a5d4faa8667569cd02ef23\" class=\"zp-Zotpress zp-Zotpress-Bib wp-block-group\">\n\n\t\t<span class=\"ZP_API_USER_ID ZP_ATTR\">1246567<\/span>\n\t\t<span class=\"ZP_ITEM_KEY ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_COLLECTION_ID ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_TAG_ID ZP_ATTR\">Umbraculida<\/span>\n\t\t<span class=\"ZP_AUTHOR ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_YEAR ZP_ATTR\"><\/span>\n        <span class=\"ZP_ITEMTYPE ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_INCLUSIVE ZP_ATTR\">1<\/span>\n\t\t<span class=\"ZP_STYLE ZP_ATTR\">eunomia-revista-en-cultura-de-la-legalidad<\/span>\n\t\t<span class=\"ZP_LIMIT ZP_ATTR\">50<\/span>\n\t\t<span class=\"ZP_SORTBY ZP_ATTR\">creator<\/span>\n\t\t<span class=\"ZP_ORDER ZP_ATTR\">asc<\/span>\n\t\t<span class=\"ZP_TITLE ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_SHOWIMAGE ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_SHOWTAGS ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_DOWNLOADABLE ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_NOTES ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_ABSTRACT ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_CITEABLE ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_TARGET ZP_ATTR\">1<\/span>\n\t\t<span class=\"ZP_URLWRAP ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_FORCENUM ZP_ATTR\"><\/span>\n        <span class=\"ZP_HIGHLIGHT ZP_ATTR\"><\/span>\n        <span class=\"ZP_POSTID ZP_ATTR\">7853<\/span>\n\t\t<span class=\"ZOTPRESS_PLUGIN_URL ZP_ATTR\">https:\/\/opistobranquis.info\/wp-content\/plugins\/zotpress\/<\/span>\n\n\t\t<div class=\"zp-List loading\">\n\t\t\t<div class=\"zp-SEO-Content\">\n\n\t\t\t<\/div><!-- .zp-zp-SEO-Content -->\n\t\t<\/div><!-- .zp-List -->\n\t<\/div><!--.zp-Zotpress-->\n\n<p><\/p>Bibliografia basada en els treballs de <b>Steve Long, 2006<\/b>. <a target=_blank href=https:\/\/opistobranquis.info\/wp-content\/uploads\/2016\/06\/Long-S.-2006-Bibliography-of-Opisthobranchia-1554-2000.pdf>Bibliography of Opisthobranchia 1554-2000<\/a> i <b>Gary McDonald, 2009<\/b>. <a href=http:\/\/escholarship.org\/uc\/item\/8115h0wz>Bibliographia Nudibranchia<\/a>, amb actualitzacions posteriors procedents d\u00b4altres fonts.<p><\/p><\/ul><p><\/p><\/p>","protected":false},"excerpt":{"rendered":"<p>Els Umbraculida s\u00f3n un petit grup de mol\u00b7luscs gaster\u00f2podes heterobranquis marins caracteritzats per una conquilla externa petita i en forma de c\u00fapula, que cobreix parcialment el dors de l&#8217;animal. S\u00f3n llimacs marins de mida petita o mitjana, amb un cos tou i una morfologia particular que els diferencia d&#8217;altres heterobranquis amb conquilla. La seva anatomia combina caracter\u00edstiques pr\u00f2pies dels gaster\u00f2podes amb altres adaptacions associades a la vida bent\u00f2nica. Els Umbraculida\u2026<\/p>\n<p class=\"continue-reading-button\"> <a class=\"continue-reading-link\" href=\"https:\/\/opistobranquis.info\/ca\/guia\/umbraculida\/\">Continua llegint<i class=\"crycon-right-dir\"><\/i><\/a><\/p>\n","protected":false},"author":291,"featured_media":4937,"parent":185,"menu_order":5055,"comment_status":"closed","ping_status":"closed","template":"","meta":{"_jetpack_hide_featured_image":false,"_jetpack_featured_image_duplicate_dismissed":0,"footnotes":""},"class_list":["post-7853","page","type-page","status-publish","has-post-thumbnail","hentry"],"jetpack_sharing_enabled":true,"_links":{"self":[{"href":"https:\/\/opistobranquis.info\/ca\/wp-json\/wp\/v2\/pages\/7853","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/opistobranquis.info\/ca\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/opistobranquis.info\/ca\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/ca\/wp-json\/wp\/v2\/users\/291"}],"replies":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/ca\/wp-json\/wp\/v2\/comments?post=7853"}],"version-history":[{"count":7,"href":"https:\/\/opistobranquis.info\/ca\/wp-json\/wp\/v2\/pages\/7853\/revisions"}],"predecessor-version":[{"id":49047,"href":"https:\/\/opistobranquis.info\/ca\/wp-json\/wp\/v2\/pages\/7853\/revisions\/49047"}],"up":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/ca\/wp-json\/wp\/v2\/pages\/185"}],"wp:featuredmedia":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/ca\/wp-json\/wp\/v2\/media\/4937"}],"wp:attachment":[{"href":"https:\/\/opistobranquis.info\/ca\/wp-json\/wp\/v2\/media?parent=7853"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}