Doris bertheloti

Doris bertheloti (d’Orbigny, 1839)

Doris bertheloti by Enric Madrenas

Class: Gastropoda  Cuvier, 1797
Subclass: Heterobranchia  J.E. Gray, 1840
Clade: Euthyneura  Spengel, 1881
Clade: Nudipleura  Wägele & Willan, 2000
Order: Nudibranchia  Cuvier, 1817
Suborder: Euctenidiacea  Tardy, 1970
Infraorder: Doridacea  Thiele, 1931
Superfamily: Doridoidea  Rafinesque, 1815
Family: Dorididae  Rafinesque, 1815
Genus: Doris  Linnaeus, 1758
Species: Doris bertheloti (d’Orbigny, 1839) [Doridigitata]


  • Doridigitata bertheloti d’Orbigny, 1839

The specimens of this species can reach nearly 40 mm in length. The color is usually brown or greenish gray with some darker spots that sometimes form two longitudinal lines running from the area of ​​the rhinophores to the gill plume. All the back of the animal is covered with several rounded tubercles of different sizes, the largest being on the center of the back, often forming ridges of aligned tubercles. Bigger tubercles are usually flanked by smaller and radially aligned tubercles, connected by smaller sized ridges. The rhinophores have a semitransparent base portion while the laminated part is yellowish or brown with numerous lamellae (30 in 200 mm specimens). The rhinophoric sheath is high and tuberculated, and its upper edge also has a pair of thick tubercles. Gill is formed by 6-7 tripinnate leaves coloured slightly lighter than the body. Gill sheath is also tuberculated along its entire surface and upper edge. The foot is light coloured and has small dark marks. The mouth has two short and stubby conical oral tentacles.

This species usually lives under rocks from the intertidal zone to the upper subtidal, where grow the encrusting sponges it feeds on. The animals, both alive and preserved, often have a rough, hard texture. The spawn consists of an undulating ribbon of about 3 mm in height, wound in a two turns spiral and about 20 mm in diameter. The eggs are white, measuring about 150 microns in diameter and are placed in vertical rows (Ortea & Bacallado, 1980).


  • Doris.  In Greek mythology, wife of Nereus, nymph of the waters and mother of the Nereids.
  • Bertheloti. Dedicated to Sabin Berthelot (1794-1880), a French naturalyst who enrolled the Navy and participated in the Napoleonic wars. He lived part of his life at the Canary islands, in his post as French consul in Tenerife. The abbreviation “Berth” is used to indicate Sabin Berthelot as an authority in the scientific botanical description and classification. He wrote the “Historia Natural de Canarias” and directed the Botanical Garden of Puerto de la Cruz.

This species is little known because it has been only registered so far in the Canary Islands, in Madeira and in few locations of the Mediterranean (Italian coast). In the Iberian Peninsula it has been reported in the coast of Granada and has recently been observed in the Catalan Costa Brava (Es Caials, L’Escala, Cala Aiguafreda, Sant Feliu de Guixols and Blanes) (VIMAR, GROC).

Known georeferenced records of the species: Doris bertheloti
: GROC 2010-2011
: Enric Madrenas
: João Pedro Silva
: Bernard Picton
: Manuel Ballesteros.
: M@re Nostrum
: Altres fonts
: Marine Regions


    Western Mediterranean:
    Eastern Mediterranean:
    Atlantic Ocean:
This chart displays the observation probability for Doris bertheloti based on our own records.

More pictures


Further reading

Cite this article as:

Ballesteros, Manuel, Enric Madrenas, Miquel Pontes et al. (2012-2017) "Doris bertheloti" in OPK-Opistobranquis, Published: 12/09/2014, Accessed: 18/12/2017 at (

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