{"id":18306,"date":"2015-02-21T16:46:25","date_gmt":"2015-02-21T15:46:25","guid":{"rendered":"https:\/\/opistobranquis.info\/?page_id=18306"},"modified":"2026-03-20T11:03:27","modified_gmt":"2026-03-20T10:03:27","slug":"roxaniella-jeffreysi","status":"publish","type":"page","link":"https:\/\/opistobranquis.info\/en\/guia\/cephalaspidea\/haminoeoidea\/roxaniella-jeffreysi\/","title":{"rendered":"Roxaniella jeffreysi"},"content":{"rendered":"<p><\/p>\n<h3><em>Roxaniella jeffreysi<\/em>\u00a0(Weinkauff, 1866)<\/h3>\n<p><span style=\"font-size:10pt;\" class=\"aligncenter\"><div id=\"featured-img-id\"><img data-recalc-dims=\"1\" decoding=\"async\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2021\/06\/FB_IMG_1623793835717.jpg?w=900&#038;ssl=1\" alt=\"Roxaniella jeffreysi @ Split, Croatia by Pero Ugarkovi\u0107\" \/><\/div><center><em>Roxaniella jeffreysi @ Split, Croatia<\/em> by Pero Ugarkovi\u0107<\/center><\/span><p><\/p><div class=\"opk-worms\"><h3>WoRMS Taxonomic Information<\/h3><div class=\"opk-success\">Current accepted name: <strong><em>Roxaniella jeffreysi<\/em> (Weinkauff, 1866)<\/strong><\/div><h4>Current classification<\/h4><table class=\"widefat striped\"><tr><th>Superdomain<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=1\">Biota<\/a><\/td><\/tr><tr><th>Kingdom<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=2\">Animalia<\/a><\/td><\/tr><tr><th>Phylum<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=51\">Mollusca<\/a><\/td><\/tr><tr><th>Class<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=101\">Gastropoda<\/a><\/td><\/tr><tr><th>Subclass<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=14712\">Heterobranchia<\/a><\/td><\/tr><tr><th>Infraclass<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=1057247\">Euthyneura<\/a><\/td><\/tr><tr><th>Subterclass<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=1057250\">Tectipleura<\/a><\/td><\/tr><tr><th>Order<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=154\">Cephalaspidea<\/a><\/td><\/tr><tr><th>Superfamily<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=411892\">Haminoeoidea<\/a><\/td><\/tr><tr><th>Family<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=23052\">Haminoeidae<\/a><\/td><\/tr><tr><th>Genus<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=754714\"><em>Roxaniella<\/em><\/a><\/td><\/tr><tr><th>Species<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=1342955\"><em>Roxaniella jeffreysi<\/em> (Weinkauff, 1866)<\/a><\/td><\/tr><\/table><p class=\"footer\">Classification provided by the World Register of Marine Species (WoRMS). [<a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=1342955\">1342955<\/a>]<\/p><\/div><strong data-start=\"3402\" data-end=\"3421\">Taxonomic note:<\/strong>\u00a0Originally described as <em data-start=\"3473\" data-end=\"3501\">Bulla (Cylichna) jeffreysi<\/em> by Weinkauff (1866), based on material previously confused with other species of the genus <em data-start=\"3593\" data-end=\"3600\">Bulla<\/em>. Subsequently, Monterosato (1884) erected the genus <em data-start=\"3653\" data-end=\"3665\">Roxaniella<\/em> for this species; however, in much of the contemporary literature it has been treated as <em data-start=\"3755\" data-end=\"3771\">Atys jeffreysi<\/em>, considering <em data-start=\"3785\" data-end=\"3797\">Roxaniella<\/em> as a synonym or subgenus of <em data-start=\"3826\" data-end=\"3832\">Atys<\/em> (Oskars <em>et al<\/em>., 2017). Recent studies based on molecular phylogeny have demonstrated that Mediterranean species traditionally assigned to <em data-start=\"3971\" data-end=\"3977\">Atys<\/em> do not form a monophyletic group, suggesting affinities with different Indo-Pacific lineages and highlighting the taxonomic instability of the group (Oskars <em>et al<\/em>., 2017, 2019). In this context, the use of <em data-start=\"4184\" data-end=\"4196\">Roxaniella<\/em> has been reinstated in recent works on the phylogeny of Haminoeidae sensu lato (Oskars <em>et al<\/em>., 2019), although its acceptance is not yet universal.<\/p>\n<p><strong>Synonyms<\/strong><\/p>\n<ul>\n<li><span id=\"aphia_ct_tu_747883\"><i>Bulla blainvilliana<\/i> R\u00e9cluz, 1843<\/span><\/li>\n<li><span id=\"aphia_ct_tu_754715\"><i>Bulla ovulata<\/i> Brocchi, 1814 <i>sensu<\/i> Jeffreys, 1856 (misidentification)<\/span><\/li>\n<li><span id=\"aphia_ct_tu_748303\"><i>Cylichna jeffreysi<\/i> Weinkauff, 1866<\/span><\/li>\n<li><em>Atys jeffreysi<\/em> (Weinkauff, 1866)<\/li>\n<\/ul>\n<p><strong>Description<br \/>\n<\/strong>Translucent animal, with the body covered by small white dots; the mantle, beneath the shell, shows irregular brown to reddish-brown blotches, also scattered over the cephalic shield and parapodial lobes (Oskars <em>et al<\/em>., 2017). The cephalic shield is short, broad, rounded, and bilobed, with well-defined small cephalic lobes; the eyes are visible through transparency. The parapodial lobes are triangular in shape (Oskars <em>et al<\/em>., 2017). The shell is translucent, whitish to yellowish in colour, cylindrical and truncated at both ends. The surface is covered by spiral striae extending over the entire shell, formed by fused rounded indentations, alternating thicker and thinner striae. Axial striae (growth lines) intersect the spiral ones, producing a characteristic decussate pattern. The aperture extends along the entire shell, narrowing towards the apex; the outer lip extends beyond the apex and arches towards the upper columella, without apical folds or torsion. The columellar lip shows a weak fold, and a small umbilical groove is present behind this fold (Oskars <em>et al<\/em>., 2017).<\/p>\n<p><strong>Biology<br \/>\n<\/strong>Specimens studied by Oskars <em>et al<\/em>. (2017) were collected between 40 and 60 m depth on Mediterranean sea bottoms, suggesting an affinity for relatively deep infra- and circalittoral habitats. No specific published information is available regarding the feeding or reproductive biology of this species.<\/p>\n<p><strong>Etymology<\/strong><\/p>\n<ul>\n<li><em>Jeffreysi<\/em>. In honor to\u00a0John Gwyn Jeffreys, (1809-1885), solicitor from Swansea, who got interested in conchology already when young. By conchology Jeffreys meant not only shell collecting, but the total science about molluscs. He began to\u00a0organize dredging expeditions, visiting the Shetlands, W. Scotland, Channel Islands, Norway etc. and accompanied by chosen specialists. He took part in some longer expeditions to W. Greenland and the Bay of Biscay and visited collecting colleagues in foreign countries. He could not come to terms with Darwinism, because, according to his view, it implied that no borders between species were to be found. &#8220;British Conchology&#8221; in 5 volumes 1862-69 is his main work. His very extensive shell collection was purchased for the Smithsonian Institution 1883 by W.H. Dall.<\/li>\n<\/ul>\n<p><strong>Distribution<\/strong><br \/>\nThe type locality of this species is La Spezia (Italy), on the coast of the Ligurian Sea (Mediterranean). The species has been documented in various areas of the Mediterranean and in the Macaronesian region (Madeira and Canary Islands) (Oskars <em>et al<\/em>., 2017; Moro <em>et al<\/em>., 2003). The specimens analysed in the redescription by Oskars <em>et al<\/em>. (2017) originate from Malta, confirming its presence in the central Mediterranean. The small size of the species and the scarcity of records may indicate that its actual distribution is insufficiently documented.<div id=\"flxmap-6a6b946f92766\" class='flxmap-container' data-flxmap='flxmap_6a6b946f92766' style=\"width:100%;height:400px;\"><\/div>\n<script data-noptimize=\"1\">\n\/* <![CDATA[ *\/\n(function(w, fn) {\n if (w.addEventListener) w.addEventListener(\"DOMContentLoaded\", fn, false);\n else if (w.attachEvent) w.attachEvent(\"onload\", fn);\n})(window, function() {\n var f = new FlexibleMap();\n f.panControl = true;\n f.mapTypeId = \"satellite\";\n f.setlocale(\"en-US\");\n f.kmlcache = \"5 minutes\";\n f.showKML(\"flxmap-6a6b946f92766\", \"https:\/\/opistobranquis.info\/wp-content\/uploads\/kms\/18306.kmz\");\n window.flxmap_6a6b946f92766 = f;\n});\n\/* ]]> *\/\n<\/script>\n<div style=\"width:100%;text-align:center;margin:12px 0 4px 0;\">\n  <div style=\"font-size:8pt;margin-bottom:6px;\">Known georeferenced records of the species: <em>Roxaniella jeffreysi<\/em><\/div>\n  <table style=\"margin:0 auto;border-collapse:collapse;font-size:8pt;\">\n    <tr><td colspan=\"2\" style=\"text-align:left;padding-bottom:4px;\"><strong>Sources:<\/strong><\/td><\/tr>\n    <tr>\n      <td style=\"vertical-align:top;padding-right:40px;\">\n        <table style=\"border-collapse:collapse;\">\n<tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/wht-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>OBIS<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/ylw-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>GROC 2010-2011<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/grn-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>Enric Madrenas<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/orange-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>Jo\u00e3o Pedro Silva<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/pink-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>Bernard Picton<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/blk-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>GBIF<\/td><\/tr>        <\/table>\n      <\/td>\n      <td style=\"vertical-align:top;\">\n        <table style=\"border-collapse:collapse;\">\n<tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/purple-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>OPK<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/red-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>VIMAR<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/blu-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>Manuel Ballesteros<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/cyan-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>M@re Nostrum<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/grey-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>Altres fonts<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/marreg.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>Marine Regions<\/td><\/tr>        <\/table>\n      <\/td>\n    <\/tr>\n  <\/table>\n<\/div>\n<\/p>\n<p><strong>Error:<\/strong> Chart.js no est\u00e1 disponible.<\/p>\n<p><strong>More pictures<\/strong><br \/>\n<div class=\"tiled-gallery type-rectangular tiled-gallery-unresized\" data-original-width=\"900\" data-carousel-extra='{&quot;blog_id&quot;:1,&quot;permalink&quot;:&quot;https:\\\/\\\/opistobranquis.info\\\/en\\\/guia\\\/cephalaspidea\\\/haminoeoidea\\\/roxaniella-jeffreysi\\\/&quot;,&quot;likes_blog_id&quot;:44192421}' itemscope itemtype=\"http:\/\/schema.org\/ImageGallery\" > <div class=\"gallery-row\" style=\"width: 900px; height: 441px;\" data-original-width=\"900\" data-original-height=\"441\" > <div class=\"gallery-group images-1\" style=\"width: 605px; height: 441px;\" data-original-width=\"605\" data-original-height=\"441\" > <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/cephalaspidea\/haminoeoidea\/roxaniella-jeffreysi\/atys-jeffreysi-gnejna-bay-mgarr-malta-by-constantino-mifsud\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"601\"> <meta itemprop=\"height\" content=\"437\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"18309\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2015\/02\/Atys-jeffreysi-@-Gnejna-Bay-Mgarr-Malta-by-Constantino-Mifsud.jpg\" data-orig-size=\"800,582\" data-comments-opened=\"\" data-image-title=\"Roxaniella jeffreysi @ Gnejna Bay, Mgarr, Malta by Constantino Mifsud\" data-image-description=\"&lt;p&gt;Roxaniella jeffreysi @ Gnejna Bay, Mgarr, Malta by Constantino Mifsud&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2015\/02\/Atys-jeffreysi-%40-Gnejna-Bay-Mgarr-Malta-by-Constantino-Mifsud.jpg?fit=300%2C218&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2015\/02\/Atys-jeffreysi-%40-Gnejna-Bay-Mgarr-Malta-by-Constantino-Mifsud.jpg?fit=800%2C582&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2015\/02\/Atys-jeffreysi-%40-Gnejna-Bay-Mgarr-Malta-by-Constantino-Mifsud.jpg?w=601&#038;h=437&#038;ssl=1\" width=\"601\" height=\"437\" loading=\"lazy\" data-original-width=\"601\" data-original-height=\"437\" itemprop=\"http:\/\/schema.org\/image\" title=\"Roxaniella jeffreysi @ Gnejna Bay, Mgarr, Malta by Constantino Mifsud\" alt=\"Roxaniella jeffreysi @ Gnejna Bay, Mgarr, Malta by Constantino Mifsud\" style=\"width: 601px; height: 437px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Roxaniella jeffreysi @ Gnejna Bay, Mgarr, Malta <\/div> <\/div> <\/div> <!-- close group --> <div class=\"gallery-group images-1\" style=\"width: 295px; height: 441px;\" data-original-width=\"295\" data-original-height=\"441\" > <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/cephalaspidea\/haminoeoidea\/roxaniella-jeffreysi\/roxaniella-jeffreysi-bahia-de-tufia-gran-canaria-26-sep-2025-by-juan-jose-calderin-penate-3\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"291\"> <meta itemprop=\"height\" content=\"437\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"48185\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2026\/03\/Roxaniella-jeffreysi-@-bahia-de-Tufia-Gran-Canaria-26-sep-2025-by-Juan-Jose-Calderin-Penate-3.jpg\" data-orig-size=\"683,1024\" data-comments-opened=\"\" data-image-title=\"Roxaniella jeffreysi @ bah\u00eda de Tufia, Gran Canaria, 26 sep 2025 by Jos\u00e9 Juan Calder\u00edn Pe\u00f1ate\" data-image-description=\"&lt;p&gt;Roxaniella jeffreysi @ bah\u00eda de Tufia, Gran Canaria, 26 sep 2025 by Jos\u00e9 Juan Calder\u00edn Pe\u00f1ate&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2026\/03\/Roxaniella-jeffreysi-%40-bahia-de-Tufia-Gran-Canaria-26-sep-2025-by-Juan-Jose-Calderin-Penate-3.jpg?fit=200%2C300&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2026\/03\/Roxaniella-jeffreysi-%40-bahia-de-Tufia-Gran-Canaria-26-sep-2025-by-Juan-Jose-Calderin-Penate-3.jpg?fit=683%2C1024&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2026\/03\/Roxaniella-jeffreysi-%40-bahia-de-Tufia-Gran-Canaria-26-sep-2025-by-Juan-Jose-Calderin-Penate-3.jpg?w=291&#038;h=437&#038;ssl=1\" width=\"291\" height=\"437\" loading=\"lazy\" data-original-width=\"291\" data-original-height=\"437\" itemprop=\"http:\/\/schema.org\/image\" title=\"Roxaniella jeffreysi @ bah\u00eda de Tufia, Gran Canaria, 26 sep 2025 by Jos\u00e9 Juan Calder\u00edn Pe\u00f1ate\" alt=\"Roxaniella jeffreysi @ bah\u00eda de Tufia, Gran Canaria, 26 sep 2025 by Jos\u00e9 Juan Calder\u00edn Pe\u00f1ate\" style=\"width: 291px; height: 437px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Roxaniella jeffreysi @ bah\u00eda de Tufia, Gran Canaria, 26 sep 2025 by Jos\u00e9 Juan Calder\u00edn Pe\u00f1ate <\/div> <\/div> <\/div> <!-- close group --> <\/div> <!-- close row --> <div class=\"gallery-row\" style=\"width: 900px; height: 682px;\" data-original-width=\"900\" data-original-height=\"682\" > <div class=\"gallery-group images-1\" style=\"width: 456px; height: 682px;\" data-original-width=\"456\" data-original-height=\"682\" > <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/cephalaspidea\/haminoeoidea\/roxaniella-jeffreysi\/roxaniella-jeffreysi-bahia-de-tufia-gran-canaria-26-sep-2025-by-juan-jose-calderin-penate-2\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"452\"> <meta itemprop=\"height\" content=\"678\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"48186\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2026\/03\/Roxaniella-jeffreysi-@-bahia-de-Tufia-Gran-Canaria-26-sep-2025-by-Juan-Jose-Calderin-Penate-2.jpg\" data-orig-size=\"417,625\" data-comments-opened=\"\" data-image-title=\"Roxaniella jeffreysi @ bah\u00eda de Tufia, Gran Canaria, 26 sep 2025 by Jos\u00e9 Juan Calder\u00edn Pe\u00f1ate\" data-image-description=\"&lt;p&gt;Roxaniella jeffreysi @ bah\u00eda de Tufia, Gran Canaria, 26 sep 2025 by Jos\u00e9 Juan Calder\u00edn Pe\u00f1ate&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2026\/03\/Roxaniella-jeffreysi-%40-bahia-de-Tufia-Gran-Canaria-26-sep-2025-by-Juan-Jose-Calderin-Penate-2.jpg?fit=200%2C300&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2026\/03\/Roxaniella-jeffreysi-%40-bahia-de-Tufia-Gran-Canaria-26-sep-2025-by-Juan-Jose-Calderin-Penate-2.jpg?fit=417%2C625&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2026\/03\/Roxaniella-jeffreysi-%40-bahia-de-Tufia-Gran-Canaria-26-sep-2025-by-Juan-Jose-Calderin-Penate-2.jpg?w=452&#038;h=678&#038;ssl=1\" width=\"452\" height=\"678\" loading=\"lazy\" data-original-width=\"452\" data-original-height=\"678\" itemprop=\"http:\/\/schema.org\/image\" title=\"Roxaniella jeffreysi @ bah\u00eda de Tufia, Gran Canaria, 26 sep 2025 by Jos\u00e9 Juan Calder\u00edn Pe\u00f1ate\" alt=\"Roxaniella jeffreysi @ bah\u00eda de Tufia, Gran Canaria, 26 sep 2025 by Jos\u00e9 Juan Calder\u00edn Pe\u00f1ate\" style=\"width: 452px; height: 678px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Roxaniella jeffreysi @ bah\u00eda de Tufia, Gran Canaria, 26 sep 2025 by Jos\u00e9 Juan Calder\u00edn Pe\u00f1ate <\/div> <\/div> <\/div> <!-- close group --> <div class=\"gallery-group images-2\" style=\"width: 444px; height: 682px;\" data-original-width=\"444\" data-original-height=\"682\" > <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/cephalaspidea\/haminoeoidea\/roxaniella-jeffreysi\/roxaniella-jeffreysi-bahia-de-tufia-gran-canaria-26-sep-2025-by-juan-jose-calderin-penate-1\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"440\"> <meta itemprop=\"height\" content=\"294\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"48187\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2026\/03\/Roxaniella-jeffreysi-@-bahia-de-Tufia-Gran-Canaria-26-sep-2025-by-Juan-Jose-Calderin-Penate-1.jpg\" data-orig-size=\"1024,684\" data-comments-opened=\"\" data-image-title=\"Roxaniella jeffreysi @ bah\u00eda de Tufia, Gran Canaria, 26 sep 2025 by Jos\u00e9 Juan Calder\u00edn Pe\u00f1ate\" data-image-description=\"&lt;p&gt;Roxaniella jeffreysi @ bah\u00eda de Tufia, Gran Canaria, 26 sep 2025 by Jos\u00e9 Juan Calder\u00edn Pe\u00f1ate&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2026\/03\/Roxaniella-jeffreysi-%40-bahia-de-Tufia-Gran-Canaria-26-sep-2025-by-Juan-Jose-Calderin-Penate-1.jpg?fit=300%2C200&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2026\/03\/Roxaniella-jeffreysi-%40-bahia-de-Tufia-Gran-Canaria-26-sep-2025-by-Juan-Jose-Calderin-Penate-1.jpg?fit=900%2C601&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2026\/03\/Roxaniella-jeffreysi-%40-bahia-de-Tufia-Gran-Canaria-26-sep-2025-by-Juan-Jose-Calderin-Penate-1.jpg?w=440&#038;h=294&#038;ssl=1\" width=\"440\" height=\"294\" loading=\"lazy\" data-original-width=\"440\" data-original-height=\"294\" itemprop=\"http:\/\/schema.org\/image\" title=\"Roxaniella jeffreysi @ bah\u00eda de Tufia, Gran Canaria, 26 sep 2025 by Jos\u00e9 Juan Calder\u00edn Pe\u00f1ate\" alt=\"Roxaniella jeffreysi @ bah\u00eda de Tufia, Gran Canaria, 26 sep 2025 by Jos\u00e9 Juan Calder\u00edn Pe\u00f1ate\" style=\"width: 440px; height: 294px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Roxaniella jeffreysi @ bah\u00eda de Tufia, Gran Canaria, 26 sep 2025 by Jos\u00e9 Juan Calder\u00edn Pe\u00f1ate <\/div> <\/div> <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/cephalaspidea\/haminoeoidea\/roxaniella-jeffreysi\/roxaniella-jeffreysi-by-oskars-et-al-2017\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"440\"> <meta itemprop=\"height\" content=\"380\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"48227\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2026\/03\/Roxaniella-jeffreysi-by-Oskars-et-al.-2017.jpg\" data-orig-size=\"888,767\" data-comments-opened=\"\" data-image-title=\"Roxaniella jeffreysi by Oskars et al. (2017)\" data-image-description=\"&lt;p&gt;Roxaniella jeffreysi by Oskars et al. (2017)&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2026\/03\/Roxaniella-jeffreysi-by-Oskars-et-al.-2017.jpg?fit=300%2C259&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2026\/03\/Roxaniella-jeffreysi-by-Oskars-et-al.-2017.jpg?fit=888%2C767&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2026\/03\/Roxaniella-jeffreysi-by-Oskars-et-al.-2017.jpg?w=440&#038;h=380&#038;ssl=1\" srcset=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2026\/03\/Roxaniella-jeffreysi-by-Oskars-et-al.-2017.jpg?w=888&amp;ssl=1 888w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2026\/03\/Roxaniella-jeffreysi-by-Oskars-et-al.-2017.jpg?resize=300%2C259&amp;ssl=1 300w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2026\/03\/Roxaniella-jeffreysi-by-Oskars-et-al.-2017.jpg?resize=150%2C130&amp;ssl=1 150w\" width=\"440\" height=\"380\" loading=\"lazy\" data-original-width=\"440\" data-original-height=\"380\" itemprop=\"http:\/\/schema.org\/image\" title=\"Roxaniella jeffreysi by Oskars et al. (2017)\" alt=\"Roxaniella jeffreysi by Oskars et al. (2017)\" style=\"width: 440px; height: 380px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Roxaniella jeffreysi by Oskars et al. (2017) <\/div> <\/div> <\/div> <!-- close group --> <\/div> <!-- close row --> <\/div><\/p>\n<p><p><strong>Bibliography<\/strong><\/p><ul><div id=\"zotpress-62cb8c7992148ecee8edcdc24bc6c921\" class=\"zp-Zotpress zp-Zotpress-Bib wp-block-group\">\n\n\t\t<span class=\"ZP_API_USER_ID ZP_ATTR\">1246567<\/span>\n\t\t<span class=\"ZP_ITEM_KEY ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_COLLECTION_ID ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_TAG_ID ZP_ATTR\">Roxaniella jeffreysi<\/span>\n\t\t<span class=\"ZP_AUTHOR ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_YEAR ZP_ATTR\"><\/span>\n        <span class=\"ZP_ITEMTYPE ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_INCLUSIVE ZP_ATTR\">1<\/span>\n\t\t<span class=\"ZP_STYLE ZP_ATTR\">eunomia-revista-en-cultura-de-la-legalidad<\/span>\n\t\t<span class=\"ZP_LIMIT ZP_ATTR\">50<\/span>\n\t\t<span class=\"ZP_SORTBY ZP_ATTR\">creator<\/span>\n\t\t<span class=\"ZP_ORDER ZP_ATTR\">asc<\/span>\n\t\t<span class=\"ZP_TITLE ZP_ATTR\"><\/span>\n\t\t<span 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Digging in the sand: new species and records of mesopsammic sea slugs in the Western Mediterranean (Mollusca: Gastropoda). <i>Journal of Natural History<\/i>, <i>59<\/i>, 5-8, 411-454. <a class=&#039;zp-DOIURL&#039; target=&#039;_blank&#039; href=&#039;https:\/\/doi.org\/10.1080\/00222933.2024.2439056&#039;>https:\/\/doi.org\/10.1080\/00222933.2024.2439056<\/a><\/div>\n<\/div>\n\t\t\t\t<\/div><!-- .zp-Entry .zpSearchResultsItem -->\t\t\t\t<div id=\"zp-ID-18306-1246567-3VWGKIQP\" data-zp-author-date='Long-2006' data-zp-date-author='2006-Long' data-zp-date='2006' data-zp-year='2006' data-zp-itemtype='book' class=\"zp-Entry zpSearchResultsItem\">\n<div class=\"csl-bib-body\" style=\"line-height: 1.35; padding-left: 1em; text-indent:-1em;\">\n  <div class=\"csl-entry\">Long, S. J. (2006). <i>Bibliography of Opisthobranchia 1554-2000. 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A molecular phylogeny of the gastropod family Haminoeidae sensu lato (Heterobranchia: Cephalaspidea): a generic revision. <i>Invertebrate Systematics 33(2) 426-472<\/i>. <a class=&#039;zp-DOIURL&#039; target=&#039;_blank&#039; href=&#039;https:\/\/doi.org\/10.1071\/IS18051&#039;>https:\/\/doi.org\/10.1071\/IS18051<\/a><\/div>\n<\/div>\n\t\t\t\t<\/div><!-- .zp-Entry .zpSearchResultsItem -->\t\t\t\t<div id=\"zp-ID-18306-1246567-7J966QCF\" data-zp-author-date='Oskars-et-al.-2017' data-zp-date-author='2017-Oskars-et-al.' data-zp-date='2017' data-zp-year='2017' data-zp-itemtype='journalArticle' class=\"zp-Entry zpSearchResultsItem\">\n<div class=\"csl-bib-body\" style=\"line-height: 1.35; padding-left: 1em; text-indent:-1em;\">\n  <div class=\"csl-entry\">Oskars, T. R., Mifsud, C., Malaquias, M. A. E. (2017). 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Marine Molluscs of the Turkish Coasts: An Updated Checklist. <i>T\u00fcbitak, Turkish Journal of Zoology 38<\/i>. <a class=&#039;zp-DOIURL&#039; target=&#039;_blank&#039; href=&#039;https:\/\/doi.org\/10.3906\/zoo-1405-78&#039;>https:\/\/doi.org\/10.3906\/zoo-1405-78<\/a><\/div>\n<\/div>\n\t\t\t\t<\/div><!-- .zp-Entry .zpSearchResultsItem -->\t\t\t\t<div id=\"zp-ID-18306-1246567-IEGCL4NB\" data-zp-author-date='Prki\u0107-et-al.-2018' data-zp-date-author='2018-Prki\u0107-et-al.' data-zp-date='2018' data-zp-year='2018' data-zp-itemtype='book' class=\"zp-Entry zpSearchResultsItem\">\n<div class=\"csl-bib-body\" style=\"line-height: 1.35; padding-left: 1em; text-indent:-1em;\">\n  <div class=\"csl-entry\">Prki\u0107, J., Petani, A., Igli\u0107, \u00d0., Lan\u010da, L. (2018). <i>Stra\u017enjo\u0161kr\u017enjaci Jadranskoga Mora: Slikovni Atlas i Popis Hrvatskih Vrsta \/ Opisthobranchs of the Adriatic Sea: Photographic Atlas and List of Croatian Species<\/i>. 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Disponible en: <a class=&#039;zp-ItemURL&#039; target=&#039;_blank&#039; href=&#039;http:\/\/www.marinespecies.org&#039;>http:\/\/www.marinespecies.org<\/a><\/div>\n<\/div>\n\t\t\t\t<\/div><!-- .zp-Entry .zpSearchResultsItem -->\n\t\t\t<\/div><!-- .zp-zp-SEO-Content -->\n\t\t<\/div><!-- .zp-List -->\n\t<\/div><!--.zp-Zotpress-->\n\n<p><\/p>Bibliography based on the works by <b>Steve Long, 2006<\/b>. <a target=_blank href=https:\/\/opistobranquis.info\/wp-content\/uploads\/2016\/06\/Long-S.-2006-Bibliography-of-Opisthobranchia-1554-2000.pdf>Bibliography of Opisthobranchia 1554-2000<\/a> and <b>Gary McDonald, 2009<\/b>. <a href=http:\/\/escholarship.org\/uc\/item\/8115h0wz>Bibliographia Nudibranchia<\/a>, with later updates from other resources.<p><\/p><\/ul><p><\/p><p><strong>Further reading<\/strong><\/p><span style=\"font-size:10pt\"><ul><ul class=\"blogroll-links\"><li><a href=\"http:\/\/www.biodiversitylibrary.org\/name\/Atys%20jeffreysi\">Biodiversity Heritage Library<\/a><\/li><li><a href=\"https:\/\/www.flickr.com\/search\/?text=Atys+jeffreysi\">Flickr pictures<\/a><\/li><li><a href=\"http:\/\/www.opistobranquis.org\/ca\/guia\/179\">GROC<\/a><\/li><li><a href=\"http:\/\/www.idscaro.net\/sci\/04_med\/class\/fam3\/species\/roxaniella_jeffreysi1.htm\">IDscaro.NET<\/a><\/li><li><a href=\"http:\/\/www.irmng.org\/aphia.php?p=taxlist&#038;tName=Atys%20jeffreysi\">Interim Register of Marine and Nonmarine Genera<\/a><\/li><li><a href=\"https:\/\/web.archive.org\/web\/20250116074557\/http:\/\/medslugs.de\/E\/Med\/roxaniella_jeffreysi\/Selection.htm\">Medslugs (Med)<\/a><\/li><li><a href=\"http:\/\/www.ncbi.nlm.nih.gov\/Taxonomy\/Browser\/wwwtax.cgi?mode=Undef&#038;name=Roxaniella%20jeffreysi&#038;lvl=0&#038;srchmode=1\">NCBI GenBank<\/a><\/li><li><a href=\"https:\/\/obis.org\/taxon\/140066\">OBIS - Search by Taxon<\/a><\/li><li><a href=\"http:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&#038;id=1342955\">World Register of Marine Species<\/a><\/li><\/ul><\/ul><\/span><p><strong>Cite this article as: <\/strong><\/p><div style=\"display: block; background: #444444; padding: 10px 30px 10px 10px; border-radius: 5px; line-height: 18px;\"><p class=\"citacio\"><b>Pontes, Miquel<\/b> (2025) <em>\"Roxaniella jeffreysi\"<\/em> in <em>OPK-Opistobranquis<\/em>. Published: 21\/02\/2015. Accessed: 30\/07\/2026. Available at (<a href=https:\/\/opistobranquis.info\/en\/?p=18306>https:\/\/opistobranquis.info\/en\/?p=18306<\/a>)<\/p><p><\/p><\/div><p><\/p><\/p>","protected":false},"excerpt":{"rendered":"<p>Roxaniella jeffreysi\u00a0(Weinkauff, 1866) Taxonomic note:\u00a0Originally described as Bulla (Cylichna) jeffreysi by Weinkauff (1866), based on material previously confused with other species of the genus Bulla. Subsequently, Monterosato (1884) erected the genus Roxaniella for this species; however, in much of the contemporary literature it has been treated as Atys jeffreysi, considering Roxaniella as a synonym or subgenus of Atys (Oskars et al., 2017). Recent studies based on molecular phylogeny have demonstrated\u2026<\/p>\n<p class=\"continue-reading-button\"> <a class=\"continue-reading-link\" href=\"https:\/\/opistobranquis.info\/en\/guia\/cephalaspidea\/haminoeoidea\/roxaniella-jeffreysi\/\">Continue reading<i class=\"crycon-right-dir\"><\/i><\/a><\/p>\n","protected":false},"author":5,"featured_media":32493,"parent":26137,"menu_order":2116,"comment_status":"closed","ping_status":"closed","template":"","meta":{"footnotes":""},"class_list":["post-18306","page","type-page","status-publish","has-post-thumbnail","hentry"],"jetpack_sharing_enabled":true,"_links":{"self":[{"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages\/18306","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/users\/5"}],"replies":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/comments?post=18306"}],"version-history":[{"count":7,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages\/18306\/revisions"}],"predecessor-version":[{"id":48226,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages\/18306\/revisions\/48226"}],"up":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages\/26137"}],"wp:featuredmedia":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/media\/32493"}],"wp:attachment":[{"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/media?parent=18306"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}