{"id":482,"date":"2012-05-16T19:08:32","date_gmt":"2012-05-16T17:08:32","guid":{"rendered":"https:\/\/opistobranquis.info\/archidoris-pseudoargusarchidoris-pseudoargusarchidoris-pseudoargus"},"modified":"2025-03-13T15:29:45","modified_gmt":"2025-03-13T14:29:45","slug":"archidoris-pseudoargus","status":"publish","type":"page","link":"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/doridina\/doridoidei\/doridoidea\/archidoris-pseudoargus\/","title":{"rendered":"Archidoris pseudoargus"},"content":{"rendered":"<p><\/p>\n<h3><em>Archidoris pseudoargus<\/em> Rapp, 1827<\/h3>\n<p><span style=\"font-size:10pt;\" class=\"aligncenter\"><div id=\"featured-img-id\"><img data-recalc-dims=\"1\" decoding=\"async\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/05\/Archidoris-pseudoargus-Eating-the-sponge-Amphilectus-fucorum-The-Burroo-Calf-of-Man-Isle-of-Man.-Photograph-%C2%A9Bernard-Picton.jpg?w=900&#038;ssl=1\" alt=\"Archidoris pseudoargus feeding on Amphilectus fucorum @ Isle of Man by Bernard Picton\" \/><\/div><center><em>Archidoris pseudoargus feeding on Amphilectus fucorum @ Isle of Man<\/em> by Bernard Picton<\/center><\/span><p><\/p><div class=\"opk-worms\"><h3>WoRMS Taxonomic Information<\/h3><div class=\"opk-success\">Current accepted name: <strong><em>Archidoris pseudoargus<\/em> (Rapp, 1827)<\/strong><\/div><h4>Current classification<\/h4><table class=\"widefat striped\"><tr><th>Superdomain<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=1\">Biota<\/a><\/td><\/tr><tr><th>Kingdom<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=2\">Animalia<\/a><\/td><\/tr><tr><th>Phylum<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=51\">Mollusca<\/a><\/td><\/tr><tr><th>Class<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=101\">Gastropoda<\/a><\/td><\/tr><tr><th>Subclass<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=14712\">Heterobranchia<\/a><\/td><\/tr><tr><th>Infraclass<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=1057247\">Euthyneura<\/a><\/td><\/tr><tr><th>Subterclass<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=1057249\">Ringipleura<\/a><\/td><\/tr><tr><th>Superorder<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=411966\">Nudipleura<\/a><\/td><\/tr><tr><th>Order<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=1762\">Nudibranchia<\/a><\/td><\/tr><tr><th>Suborder<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=1052385\">Doridina<\/a><\/td><\/tr><tr><th>Infraorder<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=246038\">Doridoidei<\/a><\/td><\/tr><tr><th>Superfamily<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=173\">Doridoidea<\/a><\/td><\/tr><tr><th>Family<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=9904\">Dorididae<\/a><\/td><\/tr><tr><th>Genus<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=137658\"><em>Archidoris<\/em><\/a><\/td><\/tr><tr><th>Species<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=138763\"><em>Archidoris pseudoargus<\/em> (Rapp, 1827)<\/a><\/td><\/tr><\/table><p class=\"footer\">Classification provided by the World Register of Marine Species (WoRMS). [<a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=138763\">138763<\/a>]<\/p><\/div><strong data-start=\"0\" data-end=\"19\">Taxonomic Note:<\/strong> The species originally described as <em data-start=\"56\" data-end=\"75\" data-is-only-node=\"\">Doris pseudoargus<\/em> Rapp, 1827 has undergone multiple taxonomic revisions over time, particularly regarding its generic placement. Historically, the name <em data-start=\"210\" data-end=\"229\">Doris tuberculata<\/em> was widely misapplied to this species due to its usage by various authors following Cuvier (1804). However, <em data-start=\"338\" data-end=\"357\">Doris pseudoargus<\/em> by Rapp (1827) represents the first valid description of the species with its distinctive morphological characteristics. A significant taxonomic change occurred when Iredale &amp; O\u2019Donoghue (1923) assigned the species to the genus <em data-start=\"586\" data-end=\"598\">Archidoris<\/em>, a decision later followed by authors such as Thompson (1966), Schmekel &amp; Portmann (1982), and Picton &amp; Morrow (1994). However, Vald\u00e9s (2002) synonymized <em data-start=\"753\" data-end=\"765\">Archidoris<\/em> with <em data-start=\"771\" data-end=\"778\">Doris<\/em>, arguing that the morphological differences were insufficient to justify a separate genus.\u00a0Recent molecular phylogenetic analyses conducted by Fern\u00e1ndez-Vilert et al. (2024) have provided new insights into the systematics of the family Discodorididae. Their results indicate that <em data-start=\"1062\" data-end=\"1069\">Doris<\/em> sensu Vald\u00e9s (2002) is not monophyletic and that <em data-start=\"1119\" data-end=\"1131\">Archidoris<\/em> should be reinstated as a valid genus. Phylogenetic results support the recognition of <em data-start=\"1219\" data-end=\"1243\">Archidoris pseudoargus<\/em> as a species that does not belong to <em data-start=\"1281\" data-end=\"1288\">Doris<\/em> but is more closely related to other northern hemisphere taxa traditionally placed within <em data-start=\"1379\" data-end=\"1391\">Archidoris<\/em> and <em data-start=\"1396\" data-end=\"1409\">Austrodoris<\/em>.<\/p>\n<p><strong>Synonyms<\/strong><\/p>\n<ul>\n<li><span id=\"aphia_ct_tu_138763\"><i>Doris pseudoargus<\/i> (Rapp, 1827)<\/span><\/li>\n<li><span id=\"aphia_ct_tu_153314\"><i>Archidoris tuberculata<\/i> (M\u00fcller, 1778) sensu Cuvier, 1804<\/span><\/li>\n<li><span id=\"aphia_ct_tu_748455\"><i>Doris areolata<\/i> Stuwitz, 1836\u00a0<\/span><\/li>\n<li><span id=\"aphia_ct_tu_153315\"><i>Doris brittanica<\/i> Johnston, 1838\u00a0<\/span><\/li>\n<li><span id=\"aphia_ct_tu_549638\"><i>Doris flavipes<\/i> Leuckart, 1828<\/span><\/li>\n<li><span id=\"aphia_ct_tu_549639\"><i>Doris leuckarti<\/i> Delle Chiaje, 1841<\/span><\/li>\n<li><span id=\"aphia_ct_tu_748479\"><i>Doris leuckartii<\/i> Delle Chiaje, 1841\u00a0<\/span><\/li>\n<li><span id=\"aphia_ct_tu_153316\"><i>Doris mera<\/i> Alder &amp; Hancock, 1844\u00a0<\/span><\/li>\n<li><span id=\"aphia_ct_tu_549640\"><i>Doris schembrii<\/i> V\u00e9rany, 1846<\/span><\/li>\n<\/ul>\n<p><strong>Description<br \/>\n<\/strong>This is one of the largest doridaceans in European waters, as it can reach up to 12 cm long, but\u00a0it is frequently found only 5-8 cm long. The body is oval and fairly flat. The color is very variable, predominantly yellowish or orange specimens with brown or reddish spots of different shades and sizes scattered throughout the notum. The mantle has many\u00a0tightly placed rounded tubercles of different sizes. Larger tubercles may have other much smaller tubercles on their surface and spicules inside. In many specimens&#8217; tubercles there are white granulations. The rhinophores have an alaminar, wide, whitish base and while the laminar\u00a0portion is yellowish and has up to 30 very thin lamellae. The rhinophoric orifice is slightly elevated and tuberculate. The gill consists of 8-10 tripinnate gill leaves paler than the body color, the uppermost gill leaves&#8217; finlets (pinnulae) may have brown or reddish-brown stains. The foot is wide and yellowish, like the underside of the mantle, which lacks all types of stains. The anterior part of the foot is furrowed and the mouth and has a pair of rounded expansions acting as oral palps.<\/p>\n<p><strong>Biology<br \/>\n<\/strong>This species of nudibranch is often found under rocks in the intertidal zones of the European Atlantic coasts. In the Mediterranean it is less common and lives in the infralittoral to a certain depth, also in commercial trawlers&#8217; fishing grounds on the upper continental shelf. It has been cited feeding on sponges like <em>Halichondria panicea<\/em>, <em>Hymeniacidon sanguinea<\/em>, <em>Myxilla incrustans<\/em>, <em>Suberites ficus<\/em> and <em>Tethya aurantia<\/em>, among others (Urgorri &amp; Besteiro, 1984; McDonald &amp; Nybakken, 2009). The spawn consists in a spirally wound ribbon with yellow or orange eggs about 150 microns in diameter (Schmekel &amp; Portmann, 1982). It is often infested\u00a0by the copepod ectoparasite <em>Doridicola agilis<\/em>.<\/p>\n<p><strong>Etymology<\/strong><\/p>\n<ul>\n<li><em>Archidoris<\/em>. From ancient Greek &#8220;archaios&#8221;, old + <em>Doris<\/em>.\u00a0In Greek mythology Doris was a sea nymph, one of the many children of Oceanus and Tethys, wife of Nereus and mother of the Nereids.<\/li>\n<li><em>Pseudoargus<\/em>. From Latin &#8220;pseudo&#8221;, false + &#8220;argus&#8221;, from the Greek <i>\u0391\u03c1\u03b3\u03bf\u03c2 (Argos)<\/i>, derived from <i>\u03b1\u03c1\u03b3\u03bf\u03c2 (argos)<\/i> meaning &#8220;glistening, shining&#8221;. In Greek myth this name belonged to both the man who built the Argo and a man with a hundred eyes. For some time it was mistaken with <em>Doris argus<\/em> Philippi, 1844 nowadays a synonym of <em>Platydoris argo<\/em>, a different species, so the specific name was changed to <em>pseudoargus,<\/em> &#8220;false argo&#8221;.<\/li>\n<\/ul>\n<p><strong>Distribution<br \/>\n<\/strong>This species has a very wide distribution as it has been quoted from northern Europe, in Norway, to the Mediterranean Sea. In the Iberian Peninsula it has been reported in almost all coastal areas and in the Balearic islands (Cervera et al. 2004). 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fullscreen; picture-in-picture; clipboard-write; encrypted-media\"><\/iframe><\/p>\n<p><iframe loading=\"lazy\" src=\"https:\/\/player.vimeo.com\/video\/784332766?dnt=1&amp;app_id=122963\" width=\"900\" height=\"506\" frameborder=\"0\" allow=\"autoplay; fullscreen; picture-in-picture; clipboard-write; encrypted-media\"><\/iframe><\/p>\n<p>&nbsp;<\/p>\n<p><strong>More pictures<br \/>\n<\/strong><div class=\"tiled-gallery type-rectangular tiled-gallery-unresized\" data-original-width=\"900\" data-carousel-extra='{&quot;blog_id&quot;:1,&quot;permalink&quot;:&quot;https:\\\/\\\/opistobranquis.info\\\/en\\\/guia\\\/nudibranchia\\\/doridina\\\/doridoidei\\\/doridoidea\\\/archidoris-pseudoargus\\\/&quot;,&quot;likes_blog_id&quot;:44192421}' itemscope itemtype=\"http:\/\/schema.org\/ImageGallery\" > <div class=\"gallery-row\" style=\"width: 900px; height: 436px;\" data-original-width=\"900\" data-original-height=\"436\" > <div class=\"gallery-group images-1\" style=\"width: 578px; height: 436px;\" data-original-width=\"578\" data-original-height=\"436\" > <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/doridina\/doridoidei\/doridoidea\/archidoris-pseudoargus\/archidoris-pseudoargus18\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"574\"> <meta itemprop=\"height\" content=\"432\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"6314\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2012\/05\/Archidoris-pseudoargus-02.jpg\" data-orig-size=\"808,608\" data-comments-opened=\"\" data-image-title=\"Archidoris pseudoargus @ Blanes, Spain by Manuel Ballesteros\" data-image-description=\"&lt;p&gt;Archidoris pseudoargus @ Blanes, Spain by Manuel Ballesteros&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/05\/Archidoris-pseudoargus-02.jpg?fit=300%2C225&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/05\/Archidoris-pseudoargus-02.jpg?fit=808%2C608&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/05\/Archidoris-pseudoargus-02.jpg?w=574&#038;h=432&#038;ssl=1\" srcset=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/05\/Archidoris-pseudoargus-02.jpg?w=808&amp;ssl=1 808w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/05\/Archidoris-pseudoargus-02.jpg?resize=300%2C225&amp;ssl=1 300w\" width=\"574\" height=\"432\" loading=\"lazy\" data-original-width=\"574\" data-original-height=\"432\" itemprop=\"http:\/\/schema.org\/image\" title=\"Archidoris pseudoargus @ Blanes, Spain by Manuel Ballesteros\" alt=\"Archidoris pseudoargus @ Blanes, Spain by Manuel Ballesteros\" style=\"width: 574px; height: 432px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Archidoris pseudoargus @ Blanes, Spain by Manuel Ballesteros <\/div> <\/div> <\/div> <!-- close group --> <div class=\"gallery-group images-2\" style=\"width: 322px; height: 436px;\" data-original-width=\"322\" data-original-height=\"436\" > <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/doridina\/doridoidei\/doridoidea\/archidoris-pseudoargus\/archidoris-pseudoargus-00-2\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"318\"> <meta itemprop=\"height\" content=\"214\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"6316\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2012\/05\/Archidoris-pseudoargus-00.jpg\" data-orig-size=\"808,544\" data-comments-opened=\"\" data-image-title=\"Archidoris pseudoargus @ Blanes, Spain by Manuel Ballesteros\" data-image-description=\"&lt;p&gt;Archidoris pseudoargus @ Blanes, Spain by Manuel Ballesteros&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/05\/Archidoris-pseudoargus-00.jpg?fit=300%2C201&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/05\/Archidoris-pseudoargus-00.jpg?fit=808%2C544&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/05\/Archidoris-pseudoargus-00.jpg?w=318&#038;h=214&#038;ssl=1\" srcset=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/05\/Archidoris-pseudoargus-00.jpg?w=808&amp;ssl=1 808w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/05\/Archidoris-pseudoargus-00.jpg?resize=300%2C201&amp;ssl=1 300w\" width=\"318\" height=\"214\" loading=\"lazy\" data-original-width=\"318\" data-original-height=\"214\" itemprop=\"http:\/\/schema.org\/image\" title=\"Archidoris pseudoargus @ Blanes, Spain by Manuel Ballesteros\" alt=\"Archidoris pseudoargus @ Blanes, Spain by Manuel Ballesteros\" style=\"width: 318px; height: 214px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Archidoris pseudoargus @ Blanes, Spain by Manuel Ballesteros <\/div> <\/div> <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/doridina\/doridoidei\/doridoidea\/archidoris-pseudoargus\/archidoris-pseudoargus-01\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"318\"> <meta itemprop=\"height\" content=\"214\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"6317\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2012\/05\/Archidoris-pseudoargus-01.jpg\" data-orig-size=\"808,544\" data-comments-opened=\"\" data-image-title=\"Archidoris pseudoargus @ Blanes, Spain by Manuel Ballesteros\" data-image-description=\"&lt;p&gt;Archidoris pseudoargus @ Blanes, Spain by Manuel Ballesteros&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/05\/Archidoris-pseudoargus-01.jpg?fit=300%2C201&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/05\/Archidoris-pseudoargus-01.jpg?fit=808%2C544&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/05\/Archidoris-pseudoargus-01.jpg?w=318&#038;h=214&#038;ssl=1\" srcset=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/05\/Archidoris-pseudoargus-01.jpg?w=808&amp;ssl=1 808w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2012\/05\/Archidoris-pseudoargus-01.jpg?resize=300%2C201&amp;ssl=1 300w\" width=\"318\" height=\"214\" loading=\"lazy\" data-original-width=\"318\" data-original-height=\"214\" itemprop=\"http:\/\/schema.org\/image\" title=\"Archidoris pseudoargus @ Blanes, Spain by Manuel Ballesteros\" alt=\"Archidoris pseudoargus @ Blanes, Spain by Manuel Ballesteros\" style=\"width: 318px; height: 214px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Archidoris pseudoargus @ Blanes, Spain by Manuel Ballesteros <\/div> <\/div> <\/div> <!-- close group --> <\/div> <!-- close row --> <div class=\"gallery-row\" style=\"width: 900px; height: 402px;\" data-original-width=\"900\" data-original-height=\"402\" > <div class=\"gallery-group images-2\" style=\"width: 300px; height: 402px;\" data-original-width=\"300\" data-original-height=\"402\" > <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/doridina\/doridoidei\/doridoidea\/archidoris-pseudoargus\/doris-pseudoargus-france-by-pascal-girard\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"296\"> <meta itemprop=\"height\" content=\"197\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"36086\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2022\/12\/Doris-pseudoargus-@-France-by-Pascal-Girard.jpg\" data-orig-size=\"1280,853\" data-comments-opened=\"\" data-image-title=\"Archidoris pseudoargus @ France by Pascal Girard\" data-image-description=\"&lt;p&gt;Archidoris pseudoargus @ France by Pascal Girard&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2022\/12\/Doris-pseudoargus-%40-France-by-Pascal-Girard.jpg?fit=300%2C200&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2022\/12\/Doris-pseudoargus-%40-France-by-Pascal-Girard.jpg?fit=900%2C599&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2022\/12\/Doris-pseudoargus-%40-France-by-Pascal-Girard.jpg?w=296&#038;h=197&#038;ssl=1\" width=\"296\" height=\"197\" loading=\"lazy\" data-original-width=\"296\" data-original-height=\"197\" itemprop=\"http:\/\/schema.org\/image\" title=\"Archidoris pseudoargus @ France by Pascal Girard\" alt=\"Archidoris pseudoargus @ France by Pascal Girard\" style=\"width: 296px; height: 197px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Archidoris pseudoargus @ France by Pascal Girard <\/div> <\/div> <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/doridina\/doridoidei\/doridoidea\/archidoris-pseudoargus\/doris-pseudoargus-by-klas-malmberg\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"296\"> <meta itemprop=\"height\" content=\"197\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"36573\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2023\/01\/Doris-pseudoargus-by-Klas-Malmberg.jpg\" data-orig-size=\"1200,797\" data-comments-opened=\"\" data-image-title=\"Archidoris pseudoargus @ Sweden by Klas Malmberg\" data-image-description=\"&lt;p&gt;Archidoris pseudoargus @ Sweden by Klas Malmberg&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/01\/Doris-pseudoargus-by-Klas-Malmberg.jpg?fit=300%2C199&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/01\/Doris-pseudoargus-by-Klas-Malmberg.jpg?fit=900%2C598&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/01\/Doris-pseudoargus-by-Klas-Malmberg.jpg?w=296&#038;h=197&#038;ssl=1\" srcset=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/01\/Doris-pseudoargus-by-Klas-Malmberg.jpg?w=1200&amp;ssl=1 1200w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/01\/Doris-pseudoargus-by-Klas-Malmberg.jpg?resize=300%2C199&amp;ssl=1 300w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/01\/Doris-pseudoargus-by-Klas-Malmberg.jpg?resize=1024%2C680&amp;ssl=1 1024w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/01\/Doris-pseudoargus-by-Klas-Malmberg.jpg?resize=150%2C100&amp;ssl=1 150w\" width=\"296\" height=\"197\" loading=\"lazy\" data-original-width=\"296\" data-original-height=\"197\" itemprop=\"http:\/\/schema.org\/image\" title=\"Archidoris pseudoargus @ Sweden by Klas Malmberg\" alt=\"Archidoris pseudoargus @ Sweden by Klas Malmberg\" style=\"width: 296px; height: 197px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Archidoris pseudoargus @ Sweden by Klas Malmberg <\/div> <\/div> <\/div> <!-- close group --> <div class=\"gallery-group images-1\" style=\"width: 600px; height: 402px;\" data-original-width=\"600\" data-original-height=\"402\" > <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/doridina\/doridoidei\/doridoidea\/archidoris-pseudoargus\/doris-pseudoargus-by-klas-malmberg-2\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"596\"> <meta itemprop=\"height\" content=\"398\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"36574\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2023\/01\/Doris-pseudoargus-by-Klas-Malmberg-2.jpg\" data-orig-size=\"1200,800\" data-comments-opened=\"\" data-image-title=\"Archidoris pseudoargus @ Sweden by Klas Malmberg\" data-image-description=\"&lt;p&gt;Archidoris pseudoargus @ Sweden by Klas Malmberg&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/01\/Doris-pseudoargus-by-Klas-Malmberg-2.jpg?fit=300%2C200&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/01\/Doris-pseudoargus-by-Klas-Malmberg-2.jpg?fit=900%2C600&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/01\/Doris-pseudoargus-by-Klas-Malmberg-2.jpg?w=596&#038;h=398&#038;ssl=1\" srcset=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/01\/Doris-pseudoargus-by-Klas-Malmberg-2.jpg?w=1200&amp;ssl=1 1200w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/01\/Doris-pseudoargus-by-Klas-Malmberg-2.jpg?resize=300%2C200&amp;ssl=1 300w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/01\/Doris-pseudoargus-by-Klas-Malmberg-2.jpg?resize=1024%2C683&amp;ssl=1 1024w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2023\/01\/Doris-pseudoargus-by-Klas-Malmberg-2.jpg?resize=150%2C100&amp;ssl=1 150w\" width=\"596\" height=\"398\" loading=\"lazy\" data-original-width=\"596\" data-original-height=\"398\" itemprop=\"http:\/\/schema.org\/image\" title=\"Archidoris pseudoargus @ Sweden by Klas Malmberg\" alt=\"Archidoris pseudoargus @ Sweden by Klas Malmberg\" style=\"width: 596px; height: 398px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Archidoris pseudoargus @ Sweden by Klas Malmberg <\/div> <\/div> <\/div> <!-- close group --> <\/div> <!-- close row --> <\/div><\/p>\n<p><p><strong>Bibliography<\/strong><\/p><ul><div id=\"zotpress-abcd49067d1a36782931c97d15df7b53\" class=\"zp-Zotpress zp-Zotpress-Bib wp-block-group\">\n\n\t\t<span class=\"ZP_API_USER_ID ZP_ATTR\">1246567<\/span>\n\t\t<span class=\"ZP_ITEM_KEY ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_COLLECTION_ID ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_TAG_ID ZP_ATTR\">Archidoris pseudoargus<\/span>\n\t\t<span class=\"ZP_AUTHOR ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_YEAR ZP_ATTR\"><\/span>\n        <span class=\"ZP_ITEMTYPE ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_INCLUSIVE ZP_ATTR\">1<\/span>\n\t\t<span class=\"ZP_STYLE ZP_ATTR\">eunomia-revista-en-cultura-de-la-legalidad<\/span>\n\t\t<span class=\"ZP_LIMIT ZP_ATTR\">50<\/span>\n\t\t<span class=\"ZP_SORTBY ZP_ATTR\">creator<\/span>\n\t\t<span class=\"ZP_ORDER ZP_ATTR\">asc<\/span>\n\t\t<span class=\"ZP_TITLE ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_SHOWIMAGE ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_SHOWTAGS ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_DOWNLOADABLE ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_NOTES ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_ABSTRACT ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_CITEABLE ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_TARGET ZP_ATTR\">1<\/span>\n\t\t<span class=\"ZP_URLWRAP ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_FORCENUM ZP_ATTR\"><\/span>\n        <span class=\"ZP_HIGHLIGHT ZP_ATTR\"><\/span>\n        <span class=\"ZP_POSTID ZP_ATTR\">482<\/span>\n\t\t<span class=\"ZOTPRESS_PLUGIN_URL ZP_ATTR\">https:\/\/opistobranquis.info\/wp-content\/plugins\/zotpress\/<\/span>\n\n\t\t<div class=\"zp-List loading\">\n\t\t\t<div class=\"zp-SEO-Content\">\n\n\t\t\t<\/div><!-- .zp-zp-SEO-Content -->\n\t\t<\/div><!-- .zp-List -->\n\t<\/div><!--.zp-Zotpress-->\n\n<p><\/p>Bibliography based on the works by <b>Steve Long, 2006<\/b>. <a target=_blank href=https:\/\/opistobranquis.info\/wp-content\/uploads\/2016\/06\/Long-S.-2006-Bibliography-of-Opisthobranchia-1554-2000.pdf>Bibliography of Opisthobranchia 1554-2000<\/a> and <b>Gary McDonald, 2009<\/b>. <a href=http:\/\/escholarship.org\/uc\/item\/8115h0wz>Bibliographia Nudibranchia<\/a>, with later updates from other resources.<p><\/p><\/ul><p><\/p><p><strong>Further reading<\/strong><\/p><span style=\"font-size:10pt\"><ul><ul class=\"blogroll-links\"><li><a href=\"http:\/\/www.asturnatura.com\/especie\/archidoris-pseudoargus.html\">AsturNatura<\/a><\/li><li><a href=\"http:\/\/www.biodiversitylibrary.org\/name\/Doris%20pseudoargus\">Biodiversity Heritage Library<\/a><\/li><li><a href=\"http:\/\/www.biologiamarina.org\/wp\/doride-antico\/\">BiologiaMarina.org<\/a><\/li><li><a href=\"http:\/\/doris.ffessm.fr\/fiche2.asp?fiche_numero=574\">DORIS<\/a><\/li><li><a href=\"https:\/\/litoraldegranada.ugr.es\/el-litoral\/el-litoral-sumergido\/fauna\/moluscos\/gasteropodos\/opistobranquios\/orden-nudibranchia\/doridaceos\/doridoideos\/doris-pseudoargus\/\">El Litoral de Granada<\/a><\/li><li><a href=\"http:\/\/www.habitas.org.uk\/marinelife\/species.asp?item=W14030#\">Encyclopedia of Marine Life of Britain and Ireland<\/a><\/li><li><a href=\"http:\/\/nature22.com\/estran22\/mollusques\/gasteropodesbis\/gasteropodes5.html\">Estran 22 - Faune et flore de la zone de balancement des mar\u00e9es en C\u00f4tes d&#039;Armor<\/a><\/li><li><a href=\"https:\/\/www.flickr.com\/photos\/56388191@N08\/sets\/72177720298227806\/\">European Sea Slugs by Morddyn<\/a><\/li><li><a href=\"https:\/\/www.flickr.com\/search\/?text=Doris%20pseudoargus\">Flickr pictures<\/a><\/li><li><a href=\"http:\/\/www.irmng.org\/aphia.php?p=taxlist&#038;tName=Doris%20pseudoargus\">Interim Register of Marine and Nonmarine Genera<\/a><\/li><li><a href=\"https:\/\/web.archive.org\/web\/20250116074557\/http:\/\/medslugs.de\/E\/Atl-NE\/Doris_pseudoargus\/Selection.htm\">Medslugs (Atl.NE)<\/a><\/li><li><a href=\"http:\/\/www.ncbi.nlm.nih.gov\/Taxonomy\/Browser\/wwwtax.cgi?mode=Undef&#038;name=Doris%20pseudoargus&#038;lvl=0&#038;srchmode=1\">NCBI GenBank<\/a><\/li><li><a href=\"http:\/\/iobis.org\/mapper\/?taxon=Doris%20pseudoargus\">OBIS - Search by Taxon<\/a><\/li><li><a href=\"http:\/\/www.nudibranch.org\/Scottish%20Nudibranchs\/doris-pseudoargus.html\">Scottish Nudibranchs<\/a><\/li><li><a href=\"http:\/\/www.seaslugforum.net\/showall\/archpseu\">Sea Slug Forum<\/a><\/li><li><a href=\"http:\/\/www.conchsoc.net\/spaccount\/Archidoris-pseudoargus\">The Conchological Society of Great Britain and Ireland<\/a><\/li><li><a href=\"http:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&#038;id=181228\">World Register of Marine Species<\/a><\/li><\/ul><\/ul><\/span><p><strong>Cite this article as: <\/strong><\/p><div style=\"display: block; background: #444444; padding: 10px 30px 10px 10px; border-radius: 5px; line-height: 18px;\"><p class=\"citacio\"><b>Pontes, Miquel<\/b> (2025) <em>\"Archidoris pseudoargus\"<\/em> in <em>OPK-Opistobranquis<\/em>. Published: 16\/05\/2012. Accessed: 04\/11\/2026. Available at (<a href=https:\/\/opistobranquis.info\/en\/?p=482>https:\/\/opistobranquis.info\/en\/?p=482<\/a>)<\/p><p><\/p><\/div><p><\/p><\/p>","protected":false},"excerpt":{"rendered":"<p>Archidoris pseudoargus Rapp, 1827 Taxonomic Note: The species originally described as Doris pseudoargus Rapp, 1827 has undergone multiple taxonomic revisions over time, particularly regarding its generic placement. Historically, the name Doris tuberculata was widely misapplied to this species due to its usage by various authors following Cuvier (1804). However, Doris pseudoargus by Rapp (1827) represents the first valid description of the species with its distinctive morphological characteristics. A significant taxonomic\u2026<\/p>\n<p class=\"continue-reading-button\"> <a class=\"continue-reading-link\" href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/doridina\/doridoidei\/doridoidea\/archidoris-pseudoargus\/\">Continue reading<i class=\"crycon-right-dir\"><\/i><\/a><\/p>\n","protected":false},"author":5,"featured_media":16140,"parent":26109,"menu_order":1,"comment_status":"closed","ping_status":"closed","template":"","meta":{"footnotes":""},"class_list":["post-482","page","type-page","status-publish","has-post-thumbnail","hentry"],"jetpack_sharing_enabled":true,"_links":{"self":[{"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages\/482","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/users\/5"}],"replies":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/comments?post=482"}],"version-history":[{"count":9,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages\/482\/revisions"}],"predecessor-version":[{"id":46227,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages\/482\/revisions\/46227"}],"up":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages\/26109"}],"wp:featuredmedia":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/media\/16140"}],"wp:attachment":[{"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/media?parent=482"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}