{"id":7687,"date":"2013-08-29T11:01:44","date_gmt":"2013-08-29T09:01:44","guid":{"rendered":"https:\/\/opistobranquis.info\/gnathodoridacea"},"modified":"2026-11-05T09:33:33","modified_gmt":"2026-11-05T08:33:33","slug":"bathydoridoidei","status":"publish","type":"page","link":"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/doridina\/bathydoridoidei\/","title":{"rendered":"Bathydoridoidei"},"content":{"rendered":"<p>Bathydoridoidei are a small group of marine nudibranchs specialised for life in deep waters. They are soft-bodied heterobranch gastropod molluscs without an external shell in the adult stage, forming a highly distinctive lineage within the dorid nudibranchs. Unlike most nudibranchs known to divers, Bathydoridoidei live in deep, generally lightless environments and are not part of the fauna observable during conventional dives.<\/p>\n<span style=\"font-size:10pt;\" class=\"aligncenter\"><div id=\"featured-img-id\"><img data-recalc-dims=\"1\" decoding=\"async\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/08\/Bathydoris_aioca_500.jpg?w=900&#038;ssl=1\" alt=\"Bathydoris aioca\" \/><\/div><center><em>Bathydoris aioca<\/em> by OPK<\/center><\/span><p><\/p>\n<p>Bathydoridoidei are associated with the deep sea and have been found at great depths, both in polar regions and in other parts of the world&#8217;s oceans. Some species of <em>Bathydoris<\/em> have been documented at depths of more than 2,000 metres, and <em>Bathydoris aioca<\/em> has been recorded at around 2,900 metres in the northeastern Pacific. These depths correspond to environments beyond the reach of sunlight, where benthic communities are subject to environmental conditions very different from those of coastal ecosystems.<\/p>\n<p>Their morphology shows several characteristics typical of dorid nudibranchs, but also features that distinguish them from other groups. The body can be relatively large and robust, with a dorsal surface often covered by small tubercles. The head bears rhinophores, which function as chemical sensory organs, while the posterior part of the body bears external branchial structures. The buccal cavity also contains strong chitinous lateral jaws, one of the characters that has contributed to distinguishing this lineage from the Doridoidei.<\/p>\n<p>The biology of Bathydoridoidei remains poorly known, largely because their habitat makes them difficult to sample. The specimens studied have mainly been obtained through dredging, oceanographic expeditions and submersibles capable of reaching great depths. In some species, remains of benthic organisms have been found in the digestive tract, indicating that they may feed on different resources available on the seafloor. In <em>Bathydoris hodgsoni<\/em>, for example, a generalist and opportunistic feeding strategy has been suggested based on the study of stomach contents.<\/p>\n<p>Life in deep and aphotic environments involves ecological conditions very different from those found in coastal ecosystems. In the absence of sunlight, photosynthetic primary production cannot directly sustain these communities, and organisms depend on organic matter reaching them from upper layers or on production associated with other processes in the deep sea. Bathydoridoidei form part of these benthic communities and represent one of the adaptations of nudibranchs to habitats far beyond the zone normally accessible to human observation.<\/p>\n<p>From an evolutionary perspective, Bathydoridoidei are particularly interesting because they constitute a deeply differentiated lineage within the nudibranchs. Anatomical and molecular studies place them as a group closely related to Doridoidei, but with their own combination of morphological characters. Their specialisation for deep environments shows that nudibranch diversification was not restricted to illuminated coastal areas, but also gave rise to lineages adapted to deep-sea communities.<\/p>\n<p>Bathydoridoidei therefore represent a very poorly known component of nudibranch diversity. Their deep-water habitat, often thousands of metres below the surface, makes them practically inaccessible to direct observation, meaning that knowledge of their biology and distribution depends mainly on oceanographic expeditions. The study of deep benthic communities and the use of submersibles continue to provide new information about this distinctive group of sea slugs.<\/p>\n<p>The taxonomy of Bathydoridoidei according to WoRMS is:<\/p>\n<p>    <style>\n        .opk-taxonomy-loader {\n            display: flex;\n            gap: 4px;\n            align-items: center;\n            justify-content: center;\n            height: 24px;\n            margin-bottom: 1em;\n        }\n        .opk-taxonomy-loader span {\n            display: block;\n            width: 4px;\n            height: 16px;\n            background: currentColor;\n            animation: opkTaxonomyBarPulse 1.2s infinite ease-in-out;\n        }\n        .opk-taxonomy-loader span:nth-child(1) { animation-delay: 0s; }\n        .opk-taxonomy-loader span:nth-child(2) { animation-delay: 0.2s; }\n        .opk-taxonomy-loader span:nth-child(3) { animation-delay: 0.4s; }\n\n        @keyframes opkTaxonomyBarPulse {\n            0%, 100% { transform: 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class=\"ZP_ITEMTYPE ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_INCLUSIVE ZP_ATTR\">1<\/span>\n\t\t<span class=\"ZP_STYLE ZP_ATTR\">eunomia-revista-en-cultura-de-la-legalidad<\/span>\n\t\t<span class=\"ZP_LIMIT ZP_ATTR\">50<\/span>\n\t\t<span class=\"ZP_SORTBY ZP_ATTR\">creator<\/span>\n\t\t<span class=\"ZP_ORDER ZP_ATTR\">asc<\/span>\n\t\t<span class=\"ZP_TITLE ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_SHOWIMAGE ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_SHOWTAGS ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_DOWNLOADABLE ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_NOTES ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_ABSTRACT ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_CITEABLE ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_TARGET ZP_ATTR\">1<\/span>\n\t\t<span class=\"ZP_URLWRAP ZP_ATTR\"><\/span>\n\t\t<span class=\"ZP_FORCENUM ZP_ATTR\"><\/span>\n        <span class=\"ZP_HIGHLIGHT ZP_ATTR\"><\/span>\n        <span class=\"ZP_POSTID ZP_ATTR\">7687<\/span>\n\t\t<span class=\"ZOTPRESS_PLUGIN_URL ZP_ATTR\">https:\/\/opistobranquis.info\/wp-content\/plugins\/zotpress\/<\/span>\n\n\t\t<div class=\"zp-List loading\">\n\t\t\t<div class=\"zp-SEO-Content\">\n\t\t\t\t<span class=\"ZP_CACHETIMER ZP_ATTR\"><\/span><span class=\"ZP_JSON 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id=\"zp-ID-7687-1246567-9HTF4T7A\" data-zp-author-date='Troncoso-et-al.-1999' data-zp-date-author='1999-Troncoso-et-al.' data-zp-date='1999' data-zp-year='1999' data-zp-itemtype='journalArticle' class=\"zp-Entry zpSearchResultsItem\">\n<div class=\"csl-bib-body\" style=\"line-height: 1.35; padding-left: 1em; text-indent:-1em;\">\n  <div class=\"csl-entry\">Troncoso, J. S., Garc\u00eda Garc\u00eda, F. J., Garc\u00eda G\u00f3mez, J. C., Ballesteros, M. (1999). Moluscos opistobranquios capturados durante las expediciones espa\u00f1olas Bentart 94 y 95, ampliando la distribuci\u00f3n de rango batim\u00e9trico de Bathydoris hodsoni. <i>Nova acta Cient\u00edfica Compostelana (Biolox\u00eda)<\/i>, <i>9<\/i>, 321-323.<\/div>\n<\/div>\n\t\t\t\t<\/div><!-- .zp-Entry .zpSearchResultsItem -->\t\t\t\t<div id=\"zp-ID-7687-1246567-6BCMIIVI\" data-zp-author-date='W\u00e4gele-et-al.-2006' data-zp-date-author='2006-W\u00e4gele-et-al.' data-zp-date='2006' data-zp-year='2006' data-zp-itemtype='journalArticle' class=\"zp-Entry zpSearchResultsItem\">\n<div class=\"csl-bib-body\" style=\"line-height: 1.35; padding-left: 1em; text-indent:-1em;\">\n  <div class=\"csl-entry\">W\u00e4gele, H., Avila, C., Ballesteros, M. (2006). Defensive glandular structures in opisthobranch molluscs: from histology to ecology. <i>Oceanography and Marine Biology<\/i>, <i>44<\/i>, 197-276.<\/div>\n<\/div>\n\t\t\t\t<\/div><!-- .zp-Entry .zpSearchResultsItem -->\n\t\t\t<\/div><!-- .zp-zp-SEO-Content -->\n\t\t<\/div><!-- .zp-List -->\n\t<\/div><!--.zp-Zotpress-->\n\n<p><\/p>Bibliography based on the works by <b>Steve Long, 2006<\/b>. <a target=_blank href=https:\/\/opistobranquis.info\/wp-content\/uploads\/2016\/06\/Long-S.-2006-Bibliography-of-Opisthobranchia-1554-2000.pdf>Bibliography of Opisthobranchia 1554-2000<\/a> and <b>Gary McDonald, 2009<\/b>. <a href=http:\/\/escholarship.org\/uc\/item\/8115h0wz>Bibliographia Nudibranchia<\/a>, with later updates from other resources.<p><\/p><\/ul><p><\/p><\/p>","protected":false},"excerpt":{"rendered":"<p>Bathydoridoidei are a small group of marine nudibranchs specialised for life in deep waters. They are soft-bodied heterobranch gastropod molluscs without an external shell in the adult stage, forming a highly distinctive lineage within the dorid nudibranchs. Unlike most nudibranchs known to divers, Bathydoridoidei live in deep, generally lightless environments and are not part of the fauna observable during conventional dives. Bathydoridoidei are associated with the deep sea and have\u2026<\/p>\n<p class=\"continue-reading-button\"> <a class=\"continue-reading-link\" href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/doridina\/bathydoridoidei\/\">Continue reading<i class=\"crycon-right-dir\"><\/i><\/a><\/p>\n","protected":false},"author":291,"featured_media":10211,"parent":7700,"menu_order":6154,"comment_status":"closed","ping_status":"closed","template":"","meta":{"_jetpack_hide_featured_image":false,"_jetpack_featured_image_duplicate_dismissed":0,"footnotes":""},"class_list":["post-7687","page","type-page","status-publish","has-post-thumbnail","hentry"],"jetpack_sharing_enabled":true,"_links":{"self":[{"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages\/7687","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/users\/291"}],"replies":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/comments?post=7687"}],"version-history":[{"count":5,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages\/7687\/revisions"}],"predecessor-version":[{"id":49056,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages\/7687\/revisions\/49056"}],"up":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages\/7700"}],"wp:featuredmedia":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/media\/10211"}],"wp:attachment":[{"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/media?parent=7687"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}