{"id":8355,"date":"2013-09-22T14:33:50","date_gmt":"2013-09-22T12:33:50","guid":{"rendered":"https:\/\/opistobranquis.info\/?page_id=8355"},"modified":"2018-02-16T11:06:59","modified_gmt":"2018-02-16T10:06:59","slug":"aeolidia-papillosa","status":"publish","type":"page","link":"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/aeolidioidea\/aeolidia-papillosa\/","title":{"rendered":"Aeolidia papillosa"},"content":{"rendered":"<p><\/p>\n<h3><em>Aeolidia papillosa<\/em> (Linnaeus, 1761)<\/h3>\n<p><span style=\"font-size:10pt;\" class=\"aligncenter\"><div id=\"featured-img-id\"><img data-recalc-dims=\"1\" decoding=\"async\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa003.jpg?w=900&#038;ssl=1\" alt=\"Aeolidia papillosa by Manuel Ballesteros\" \/><\/div><center><em>Aeolidia papillosa<\/em> by Manuel Ballesteros<\/center><\/span><p><\/p><div class=\"opk-worms\"><h3>WoRMS Taxonomic Information<\/h3><div class=\"opk-success\">Current accepted name: <strong><em>Aeolidia papillosa<\/em> (Linnaeus, 1761)<\/strong><\/div><h4>Current classification<\/h4><table class=\"widefat striped\"><tr><th>Superdomain<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=1\">Biota<\/a><\/td><\/tr><tr><th>Kingdom<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=2\">Animalia<\/a><\/td><\/tr><tr><th>Phylum<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=51\">Mollusca<\/a><\/td><\/tr><tr><th>Class<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=101\">Gastropoda<\/a><\/td><\/tr><tr><th>Subclass<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=14712\">Heterobranchia<\/a><\/td><\/tr><tr><th>Infraclass<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=1057247\">Euthyneura<\/a><\/td><\/tr><tr><th>Subterclass<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=1057249\">Ringipleura<\/a><\/td><\/tr><tr><th>Superorder<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=411966\">Nudipleura<\/a><\/td><\/tr><tr><th>Order<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=1762\">Nudibranchia<\/a><\/td><\/tr><tr><th>Suborder<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=1565561\">Aeolidina<\/a><\/td><\/tr><tr><th>Superfamily<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=189\">Aeolidioidea<\/a><\/td><\/tr><tr><th>Family<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=192\">Aeolidiidae<\/a><\/td><\/tr><tr><th>Genus<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=137630\"><em>Aeolidia<\/em><\/a><\/td><\/tr><tr><th>Species<\/th><td><a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=138709\"><em>Aeolidia papillosa<\/em> (Linnaeus, 1761)<\/a><\/td><\/tr><\/table><p class=\"footer\">Classification provided by the World Register of Marine Species (WoRMS). [<a target=\"_blank\" href=\"https:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&id=138709\">138709<\/a>]<\/p><\/div><strong>Synonyms<\/strong><\/p>\n<ul>\n<li><em>Aeolidia herculea<\/em> Troschel, 1866<\/li>\n<li><em>Aeolidia papillosa var. pacifica<\/em> Bergh, 1879<\/li>\n<li><em>Aeolidia serotina<\/em> Bergh, 1873<\/li>\n<li><em>Aeolis lesliana<\/em> MacGillivray, 1843<\/li>\n<li><em>Aeolis murrayana<\/em> MacGillivray, 1843<\/li>\n<li><em>Doris bodoensis<\/em> Gunnerus, 1770<\/li>\n<li><em>Doris vermigera<\/em> Turton, 1807<\/li>\n<li><em>Eolidia zetlandica<\/em> Forbes &amp; Goodsir, 1839<\/li>\n<li><em>Eolis campbellii<\/em> Cunningham, 1871<\/li>\n<li><em>Eolis cuvieri<\/em> Lamarck, 1819<\/li>\n<li><em>Eolis farinacea<\/em> Stimpson, 1853<\/li>\n<li><em>Eolis obtusalis<\/em> Alder &amp; Hancock, 1842<\/li>\n<li><em>Eolis papillosa<\/em> (Linnaeus, 1761)<\/li>\n<li><em>Eolis papillosa<\/em> var. albina Dautzenberg &amp; Durouchoux, 1913<\/li>\n<li><em>Eolis plumata<\/em> Dalyell, 1853<\/li>\n<li><em>Eolis rosea<\/em> Alder &amp; Hancock, 1842<\/li>\n<li><strong><em>Limax papillosus<\/em> Linnaeus, 1761 (original)<\/strong><\/li>\n<\/ul>\n<p><strong>Description<br \/>\n<\/strong>This is the largest aeolidacean of the European coasts, where the largest specimens seem to reach up to\u00a012 cm in length, although they are usually about half this size. The body is wide and flat, quite opaque and with a highly variable coloration, with specimens\u00a0ranging from white to\u00a0gray to yellowish brown or even purple as the base color, with many spots of white or gray or brown or even dark violet; according to their density, they change the animal&#8217;s overall appearance. These spots are more numerous in large specimens. In the front of the head it usually has a &#8220;V&#8221;\u00a0shaped stain, it&#8217;s base between the rhinophores and with the arms extended to\u00a0the oral tentacles&#8217; upper side. This stain can also extend to a rhombus or crescent shape over the pericardial area, located behind the rhinophores and not usually be covered by the cerata. The anal papilla, usually white, is hidden under the cerata. The cerata, somewhat darker than the body color, are\u00a0pointed, often have a white apex and are very numerous, as they are arranged in up to 25 transverse rows of 12-24 cerata per row on each side of the body, they have a somewhat flattened shape and they tend to be &#8220;combed&#8221; backwards. The first cerata are smaller than the rest and are located well\u00a0ahead of the rhinophores. Smooth, short and straight, the rhinophores have a conical shape with a truncated tip, and they are contractile. Like the cerata, the rhinophores have light colored apex, whitish or yellowish, and a base color similar or somewhat darker than the body, with a darker shade when contracted. Oral tentacles are somewhat longer than the rhinophores, and are separated by a distance equivalent to three times the base of one of them. The eyes are internal and are located at the base of the rhinophores, although they are usually not visible. The foot is quite wide, especially in the front, where a pair of\u00a0characteristic propodial tentacles are found. The sole is translucent white and allows to see some pinkish internal organs (genitalia) by transparency. Anatomically it is very similar to <em>A. filomenae<\/em>, but there are some differences: The cerata of <em>A. filomenae<\/em> are flattened, slightly hook-shaped and usually have a paler shade than\u00a0the rest of the body while the cerata <em>A. papillosa<\/em> are usually darker and thinner.<\/p>\n<p><strong>Biology<br \/>\n<\/strong>As the species <em>Aeolidia filomenae<\/em>, which it has traditionally been confused with, it feeds on sea anemones. In the intertidal and shallow water, even in brackish water, it usually feeds on <em>Actinia equina<\/em>, but also on <em>Actinia fragacea, Anemonia viridis<\/em> and <em>Metridium senile<\/em>. In the sublittoral it is known to feed on <em>Actinothoe sphyrodeta, Aiptasia couchi, Alcyonium digitatum, Anemonia, Sagartia, Sagartiogeton, Anthopleura ballii, Bunodactis, Cereus pedunculatus, Corynactis viridis, Diadumene<\/em> and <em>Tealia<\/em>. It has also been found down to 800 meters deep. Before attacking its preys, it widely extends the sensory tentacles and secretes abundant mucus to protect itself against the stinging filaments projected by the anemone\u00a0when it feels in danger. It is known that, like other aeolidaceans, it is able to store\u00a0the active\u00a0nematocysts from the eaten anemones and using them as a defense by passing them to the tips of the cerata. When it feels attacked it releases the accumulated nematocysts, escaping from a dog&#8217;s\u00a0jaws (<a href=\"http:\/\/www.conchsoc.net\/spAccount\/aeolidia-papillosa\">I.Smith, Conchological Society<\/a>), or repelling\u00a0a starfish, that avoided touching it at all ( <a href=\"http:\/\/www.seaslugforum.net\/find\/19418\">J.Kocian, 11\/07\/2007, Sea Slug Forum<\/a>). Spawn, usually laid\u00a0on hard substrates between January and August, consists of a string a few millimeters in diameter with capsules containing small white eggs (occasionally pink) arranged in a spiral with a characteristic zigzag. The larvae are dispersed in the plankton after hatching, but smaller juveniles are not usually found on the coasts, as they may live in greater depths.<\/p>\n<p><strong>Etymology<\/strong><\/p>\n<ul>\n<li><em>Aeolidia<\/em> from Aeolis, Greek god of the wind<\/li>\n<li><em>Papillosa<\/em> with papillae, short processes<\/li>\n<\/ul>\n<p><strong>Distribution<br \/>\n<\/strong>In the European Atlantic coasts it is found from the White Sea to Portugal, passing through the North Sea (in Dutch waters it is one of the most common species, with densities of up to 10 individuals per square meter), the British Isles, English Channel\u00a0and the French and Spanish Atlantic coast, although it seems to show preference for the colder northern waters. It is also found on the coasts of North America, both the Atlantic and the Pacific, from Alaska to California. The study of <em>Aeolidia papillosa<\/em> compared with the very similar <em>Aeolidia filomenae<\/em> reveals that samples from different localities of the Atlantic coast of Europe, attributed so far to the first species actually belong to the second. The accepted variability in the color pattern of the first species masks the existence of a second European pseudocryptic species of this genus.<br \/>\n<div id=\"flxmap-6a6768f52a09e\" class='flxmap-container' data-flxmap='flxmap_6a6768f52a09e' style=\"width:100%;height:400px;\"><\/div>\n<script data-noptimize=\"1\">\n\/* <![CDATA[ *\/\n(function(w, fn) {\n if (w.addEventListener) w.addEventListener(\"DOMContentLoaded\", fn, false);\n else if (w.attachEvent) w.attachEvent(\"onload\", fn);\n})(window, function() {\n var f = new FlexibleMap();\n f.panControl = true;\n f.mapTypeId = \"satellite\";\n f.setlocale(\"en-US\");\n f.kmlcache = \"5 minutes\";\n f.showKML(\"flxmap-6a6768f52a09e\", \"https:\/\/opistobranquis.info\/wp-content\/uploads\/kms\/8355.kmz\");\n window.flxmap_6a6768f52a09e = f;\n});\n\/* ]]> *\/\n<\/script>\n<div style=\"width:100%;text-align:center;margin:12px 0 4px 0;\">\n  <div style=\"font-size:8pt;margin-bottom:6px;\">Known georeferenced records of the species: <em>Aeolidia papillosa<\/em><\/div>\n  <table style=\"margin:0 auto;border-collapse:collapse;font-size:8pt;\">\n    <tr><td colspan=\"2\" style=\"text-align:left;padding-bottom:4px;\"><strong>Sources:<\/strong><\/td><\/tr>\n    <tr>\n      <td style=\"vertical-align:top;padding-right:40px;\">\n        <table style=\"border-collapse:collapse;\">\n<tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/wht-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>OBIS<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/ylw-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>GROC 2010-2011<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/grn-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>Enric Madrenas<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/orange-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>Jo\u00e3o Pedro Silva<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/pink-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>Bernard Picton<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/blk-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>GBIF<\/td><\/tr>        <\/table>\n      <\/td>\n      <td style=\"vertical-align:top;\">\n        <table style=\"border-collapse:collapse;\">\n<tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/purple-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>OPK<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/red-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>VIMAR<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/blu-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>Manuel Ballesteros<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/cyan-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>M@re Nostrum<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/grey-circle-lv.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>Altres fonts<\/td><\/tr><tr><td style='padding:1px 6px 1px 0;vertical-align:middle'><img data-recalc-dims=\"1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/kms\/marreg.png?w=900&#038;ssl=1\" style='display:block'><\/td><td style='padding:1px 0;vertical-align:middle;white-space:nowrap'>Marine Regions<\/td><\/tr>        <\/table>\n      <\/td>\n    <\/tr>\n  <\/table>\n<\/div>\n<\/p>\n<p><strong>Error:<\/strong> Chart.js no est\u00e1 disponible.<\/p>\n<p><strong>More pictures<\/strong><br \/>\n<div class=\"tiled-gallery type-rectangular tiled-gallery-unresized\" data-original-width=\"900\" data-carousel-extra='{&quot;blog_id&quot;:1,&quot;permalink&quot;:&quot;https:\\\/\\\/opistobranquis.info\\\/en\\\/guia\\\/nudibranchia\\\/aeolidioidea\\\/aeolidia-papillosa\\\/&quot;,&quot;likes_blog_id&quot;:44192421}' itemscope itemtype=\"http:\/\/schema.org\/ImageGallery\" > <div class=\"gallery-row\" style=\"width: 900px; height: 433px;\" data-original-width=\"900\" data-original-height=\"433\" > <div class=\"gallery-group images-2\" style=\"width: 287px; height: 433px;\" data-original-width=\"287\" data-original-height=\"433\" > <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/aeolidioidea\/aeolidia-papillosa\/aeolidia-papillosa-spawn-coil-skerries-portrush-co-antrim-northern-ireland-15m-30-april-2007-photograph-bernard-picton\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"283\"> <meta itemprop=\"height\" content=\"213\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"16135\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-Spawn-coil.-Skerries-Portrush-Co-Antrim-Northern-Ireland.-15m.-30-April-2007.-Photograph-\u00a9Bernard-Picton.jpg\" data-orig-size=\"800,600\" data-comments-opened=\"\" data-image-title=\"Aeolidia papillosa Spawn coil. Skerries, Portrush, Co Antrim, Northern Ireland. 15m. 30 April 2007. &amp;#8211; Photograph \u00a9Bernard Picton\" data-image-description=\"&lt;p&gt;Aeolidia papillosa Spawn coil. Skerries, Portrush, Co Antrim, Northern Ireland. 15m. 30 April 2007. &amp;#8211; Photograph \u00a9Bernard Picton&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-Spawn-coil.-Skerries-Portrush-Co-Antrim-Northern-Ireland.-15m.-30-April-2007.-Photograph-%C2%A9Bernard-Picton.jpg?fit=300%2C225&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-Spawn-coil.-Skerries-Portrush-Co-Antrim-Northern-Ireland.-15m.-30-April-2007.-Photograph-%C2%A9Bernard-Picton.jpg?fit=800%2C600&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-Spawn-coil.-Skerries-Portrush-Co-Antrim-Northern-Ireland.-15m.-30-April-2007.-Photograph-%C2%A9Bernard-Picton.jpg?w=283&#038;h=213&#038;ssl=1\" width=\"283\" height=\"213\" loading=\"lazy\" data-original-width=\"283\" data-original-height=\"213\" itemprop=\"http:\/\/schema.org\/image\" title=\"Aeolidia papillosa Spawn coil. Skerries, Portrush, Co Antrim, Northern Ireland. 15m. 30 April 2007. - Photograph \u00a9Bernard Picton\" alt=\"Aeolidia papillosa Spawn coil. Skerries, Portrush, Co Antrim, Northern Ireland. 15m. 30 April 2007. - Photograph \u00a9Bernard Picton\" style=\"width: 283px; height: 213px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Aeolidia papillosa Spawn coil. Skerries, Portrush, Co Antrim, Northern Ireland. 15m. 30 April 2007. &#8211; Photograph \u00a9Bernard Picton <\/div> <\/div> <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/aeolidioidea\/aeolidia-papillosa\/aeolidia-papillosa-photograph-bernard-picton\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"283\"> <meta itemprop=\"height\" content=\"212\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"16136\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-Photograph-\u00a9Bernard-Picton.jpg\" data-orig-size=\"800,600\" data-comments-opened=\"\" data-image-title=\"Aeolidia papillosa &amp;#8211; Photograph \u00a9Bernard Picton\" data-image-description=\"&lt;p&gt;Aeolidia papillosa &amp;#8211; Photograph \u00a9Bernard Picton&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-Photograph-%C2%A9Bernard-Picton.jpg?fit=300%2C225&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-Photograph-%C2%A9Bernard-Picton.jpg?fit=800%2C600&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-Photograph-%C2%A9Bernard-Picton.jpg?w=283&#038;h=212&#038;ssl=1\" width=\"283\" height=\"212\" loading=\"lazy\" data-original-width=\"283\" data-original-height=\"212\" itemprop=\"http:\/\/schema.org\/image\" title=\"Aeolidia papillosa - Photograph \u00a9Bernard Picton\" alt=\"Aeolidia papillosa - Photograph \u00a9Bernard Picton\" style=\"width: 283px; height: 212px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Aeolidia papillosa <\/div> <\/div> <\/div> <!-- close group --> <div class=\"gallery-group images-1\" style=\"width: 290px; height: 433px;\" data-original-width=\"290\" data-original-height=\"433\" > <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/aeolidioidea\/aeolidia-papillosa\/aeolidia-papillosa-by-alexander-semenov\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"286\"> <meta itemprop=\"height\" content=\"429\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"17071\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-by-Alexander-Semenov.jpg\" data-orig-size=\"600,900\" data-comments-opened=\"\" data-image-title=\"Aeolidia papillosa by Alexander Semenov\" data-image-description=\"&lt;p&gt;Aeolidia papillosa by Alexander Semenov&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-by-Alexander-Semenov.jpg?fit=200%2C300&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-by-Alexander-Semenov.jpg?fit=600%2C900&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-by-Alexander-Semenov.jpg?w=286&#038;h=429&#038;ssl=1\" srcset=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-by-Alexander-Semenov.jpg?w=600&amp;ssl=1 600w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-by-Alexander-Semenov.jpg?resize=100%2C150&amp;ssl=1 100w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-by-Alexander-Semenov.jpg?resize=200%2C300&amp;ssl=1 200w\" width=\"286\" height=\"429\" loading=\"lazy\" data-original-width=\"286\" data-original-height=\"429\" itemprop=\"http:\/\/schema.org\/image\" title=\"Aeolidia papillosa by Alexander Semenov\" alt=\"Aeolidia papillosa by Alexander Semenov\" style=\"width: 286px; height: 429px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Aeolidia papillosa by Alexander Semenov <\/div> <\/div> <\/div> <!-- close group --> <div class=\"gallery-group images-2\" style=\"width: 323px; height: 433px;\" data-original-width=\"323\" data-original-height=\"433\" > <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/aeolidioidea\/aeolidia-papillosa\/drama-aeolidia-papillosa-about-to-eat-another-nudibranch-by-alexander-semenov\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"319\"> <meta itemprop=\"height\" content=\"212\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"17098\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Drama-Aeolidia-papillosa-about-to-eat-another-nudibranch-by-Alexander-Semenov.jpg\" data-orig-size=\"1024,683\" data-comments-opened=\"\" data-image-title=\"Drama: Aeolidia papillosa about to eat another nudibranch by Alexander Semenov\" data-image-description=\"&lt;p&gt;Drama: Aeolidia papillosa about to eat another nudibranch by Alexander Semenov&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Drama-Aeolidia-papillosa-about-to-eat-another-nudibranch-by-Alexander-Semenov.jpg?fit=300%2C200&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Drama-Aeolidia-papillosa-about-to-eat-another-nudibranch-by-Alexander-Semenov.jpg?fit=900%2C600&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Drama-Aeolidia-papillosa-about-to-eat-another-nudibranch-by-Alexander-Semenov.jpg?w=319&#038;h=212&#038;ssl=1\" srcset=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Drama-Aeolidia-papillosa-about-to-eat-another-nudibranch-by-Alexander-Semenov.jpg?w=1024&amp;ssl=1 1024w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Drama-Aeolidia-papillosa-about-to-eat-another-nudibranch-by-Alexander-Semenov.jpg?resize=150%2C100&amp;ssl=1 150w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Drama-Aeolidia-papillosa-about-to-eat-another-nudibranch-by-Alexander-Semenov.jpg?resize=300%2C200&amp;ssl=1 300w\" width=\"319\" height=\"212\" loading=\"lazy\" data-original-width=\"319\" data-original-height=\"212\" itemprop=\"http:\/\/schema.org\/image\" title=\"Drama: Aeolidia papillosa about to eat another nudibranch by Alexander Semenov\" alt=\"Drama: Aeolidia papillosa about to eat another nudibranch by Alexander Semenov\" style=\"width: 319px; height: 212px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Drama: Aeolidia papillosa about to eat another nudibranch by Alexander Semenov <\/div> <\/div> <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/aeolidioidea\/aeolidia-papillosa\/aeolidia-papillosa-1-by-alexander-semenov\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"319\"> <meta itemprop=\"height\" content=\"213\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"17117\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-1-by-Alexander-Semenov.jpg\" data-orig-size=\"880,587\" data-comments-opened=\"\" data-image-title=\"Aeolidia papillosa by Alexander Semenov\" data-image-description=\"&lt;p&gt;Aeolidia papillosa by Alexander Semenov&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-1-by-Alexander-Semenov.jpg?fit=300%2C200&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-1-by-Alexander-Semenov.jpg?fit=880%2C587&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-1-by-Alexander-Semenov.jpg?w=319&#038;h=213&#038;ssl=1\" srcset=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-1-by-Alexander-Semenov.jpg?w=880&amp;ssl=1 880w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-1-by-Alexander-Semenov.jpg?resize=150%2C100&amp;ssl=1 150w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-1-by-Alexander-Semenov.jpg?resize=300%2C200&amp;ssl=1 300w\" width=\"319\" height=\"213\" loading=\"lazy\" data-original-width=\"319\" data-original-height=\"213\" itemprop=\"http:\/\/schema.org\/image\" title=\"Aeolidia papillosa by Alexander Semenov\" alt=\"Aeolidia papillosa by Alexander Semenov\" style=\"width: 319px; height: 213px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Aeolidia papillosa by Alexander Semenov <\/div> <\/div> <\/div> <!-- close group --> <\/div> <!-- close row --> <div class=\"gallery-row\" style=\"width: 900px; height: 417px;\" data-original-width=\"900\" data-original-height=\"417\" > <div class=\"gallery-group images-1\" style=\"width: 623px; height: 417px;\" data-original-width=\"623\" data-original-height=\"417\" > <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/aeolidioidea\/aeolidia-papillosa\/aeolidia-papillosa-2-by-alexander-semenov\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"619\"> <meta itemprop=\"height\" content=\"413\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"17118\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-2-by-Alexander-Semenov.jpg\" data-orig-size=\"880,587\" data-comments-opened=\"\" data-image-title=\"Aeolidia papillosa by Alexander Semenov\" data-image-description=\"&lt;p&gt;Aeolidia papillosa by Alexander Semenov&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-2-by-Alexander-Semenov.jpg?fit=300%2C200&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-2-by-Alexander-Semenov.jpg?fit=880%2C587&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-2-by-Alexander-Semenov.jpg?w=619&#038;h=413&#038;ssl=1\" srcset=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-2-by-Alexander-Semenov.jpg?w=880&amp;ssl=1 880w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-2-by-Alexander-Semenov.jpg?resize=150%2C100&amp;ssl=1 150w, https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2014\/12\/Aeolidia-papillosa-2-by-Alexander-Semenov.jpg?resize=300%2C200&amp;ssl=1 300w\" width=\"619\" height=\"413\" loading=\"lazy\" data-original-width=\"619\" data-original-height=\"413\" itemprop=\"http:\/\/schema.org\/image\" title=\"Aeolidia papillosa by Alexander Semenov\" alt=\"Aeolidia papillosa by Alexander Semenov\" style=\"width: 619px; height: 413px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Aeolidia papillosa by Alexander Semenov <\/div> <\/div> <\/div> <!-- close group --> <div class=\"gallery-group images-2\" style=\"width: 277px; height: 417px;\" data-original-width=\"277\" data-original-height=\"417\" > <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/aeolidioidea\/aeolidia-papillosa\/olympus-digital-camera-97\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"273\"> <meta itemprop=\"height\" content=\"205\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"23410\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-@-Netherlands-20-02-2017-by-Stefan-Verheyen.jpg\" data-orig-size=\"2304,1728\" data-comments-opened=\"\" data-image-title=\"Aeolidia papillosa @ Netherlands 20-02-2017 by Stefan Verheyen\" data-image-description=\"&lt;p&gt;Aeolidia papillosa @ Netherlands 20-02-2017 by Stefan Verheyen&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-%40-Netherlands-20-02-2017-by-Stefan-Verheyen.jpg?fit=300%2C225&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-%40-Netherlands-20-02-2017-by-Stefan-Verheyen.jpg?fit=900%2C675&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-%40-Netherlands-20-02-2017-by-Stefan-Verheyen.jpg?w=273&#038;h=205&#038;ssl=1\" width=\"273\" height=\"205\" loading=\"lazy\" data-original-width=\"273\" data-original-height=\"205\" itemprop=\"http:\/\/schema.org\/image\" title=\"Aeolidia papillosa @ Netherlands 20-02-2017 by Stefan Verheyen\" alt=\"Aeolidia papillosa @ Netherlands 20-02-2017 by Stefan Verheyen\" style=\"width: 273px; height: 205px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Aeolidia papillosa @ Netherlands 20-02-2017 by Stefan Verheyen <\/div> <\/div> <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/aeolidioidea\/aeolidia-papillosa\/olympus-digital-camera-98\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"273\"> <meta itemprop=\"height\" content=\"204\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"23411\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-@-Netherlands-20-02-2017-by-Stefan-Verheyen-2.jpg\" data-orig-size=\"2304,1728\" data-comments-opened=\"\" data-image-title=\"Aeolidia papillosa @ Netherlands 20-02-2017 by Stefan Verheyen\" data-image-description=\"&lt;p&gt;Aeolidia papillosa @ Netherlands 20-02-2017 by Stefan Verheyen&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-%40-Netherlands-20-02-2017-by-Stefan-Verheyen-2.jpg?fit=300%2C225&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-%40-Netherlands-20-02-2017-by-Stefan-Verheyen-2.jpg?fit=900%2C675&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-%40-Netherlands-20-02-2017-by-Stefan-Verheyen-2.jpg?w=273&#038;h=204&#038;ssl=1\" width=\"273\" height=\"204\" loading=\"lazy\" data-original-width=\"273\" data-original-height=\"204\" itemprop=\"http:\/\/schema.org\/image\" title=\"Aeolidia papillosa @ Netherlands 20-02-2017 by Stefan Verheyen\" alt=\"Aeolidia papillosa @ Netherlands 20-02-2017 by Stefan Verheyen\" style=\"width: 273px; height: 204px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Aeolidia papillosa @ Netherlands 20-02-2017 by Stefan Verheyen <\/div> <\/div> <\/div> <!-- close group --> <\/div> <!-- close row --> <div class=\"gallery-row\" style=\"width: 900px; height: 226px;\" data-original-width=\"900\" data-original-height=\"226\" > <div class=\"gallery-group images-1\" style=\"width: 300px; height: 226px;\" data-original-width=\"300\" data-original-height=\"226\" > <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/aeolidioidea\/aeolidia-papillosa\/olympus-digital-camera-99\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"296\"> <meta itemprop=\"height\" content=\"222\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"23412\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-@-Netherlands-7-02-2017-by-Stefan-Verheyen.jpg\" data-orig-size=\"2048,1536\" data-comments-opened=\"\" data-image-title=\"Aeolidia papillosa @ Netherlands 7-02-2017 by Stefan Verheyen\" data-image-description=\"&lt;p&gt;Aeolidia papillosa @ Netherlands 7-02-2017 by Stefan Verheyen&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-%40-Netherlands-7-02-2017-by-Stefan-Verheyen.jpg?fit=300%2C225&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-%40-Netherlands-7-02-2017-by-Stefan-Verheyen.jpg?fit=900%2C675&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-%40-Netherlands-7-02-2017-by-Stefan-Verheyen.jpg?w=296&#038;h=222&#038;ssl=1\" width=\"296\" height=\"222\" loading=\"lazy\" data-original-width=\"296\" data-original-height=\"222\" itemprop=\"http:\/\/schema.org\/image\" title=\"Aeolidia papillosa @ Netherlands 7-02-2017 by Stefan Verheyen\" alt=\"Aeolidia papillosa @ Netherlands 7-02-2017 by Stefan Verheyen\" style=\"width: 296px; height: 222px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Aeolidia papillosa @ Netherlands 7-02-2017 by Stefan Verheyen <\/div> <\/div> <\/div> <!-- close group --> <div class=\"gallery-group images-1\" style=\"width: 300px; height: 226px;\" data-original-width=\"300\" data-original-height=\"226\" > <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/aeolidioidea\/aeolidia-papillosa\/olympus-digital-camera-111\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"296\"> <meta itemprop=\"height\" content=\"222\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"23440\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-@-Stavenisse-Netherlands-31-01-2017-by-Stefan-Verheyen.jpg\" data-orig-size=\"2140,1606\" data-comments-opened=\"\" data-image-title=\"Aeolidia papillosa @ Stavenisse, Netherlands 31-01-2017 by Stefan Verheyen\" data-image-description=\"&lt;p&gt;Aeolidia papillosa @ Stavenisse, Netherlands 31-01-2017 by Stefan Verheyen&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-%40-Stavenisse-Netherlands-31-01-2017-by-Stefan-Verheyen.jpg?fit=300%2C225&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-%40-Stavenisse-Netherlands-31-01-2017-by-Stefan-Verheyen.jpg?fit=900%2C675&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2013\/09\/Aeolidia-papillosa-%40-Stavenisse-Netherlands-31-01-2017-by-Stefan-Verheyen.jpg?w=296&#038;h=222&#038;ssl=1\" width=\"296\" height=\"222\" loading=\"lazy\" data-original-width=\"296\" data-original-height=\"222\" itemprop=\"http:\/\/schema.org\/image\" title=\"Aeolidia papillosa @ Stavenisse, Netherlands 31-01-2017 by Stefan Verheyen\" alt=\"Aeolidia papillosa @ Stavenisse, Netherlands 31-01-2017 by Stefan Verheyen\" style=\"width: 296px; height: 222px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Aeolidia papillosa @ Stavenisse, Netherlands 31-01-2017 by Stefan Verheyen <\/div> <\/div> <\/div> <!-- close group --> <div class=\"gallery-group images-1\" style=\"width: 300px; height: 226px;\" data-original-width=\"300\" data-original-height=\"226\" > <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/aeolidioidea\/aeolidia-papillosa\/olympus-digital-camera-135\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"296\"> <meta itemprop=\"height\" content=\"222\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"24240\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2017\/12\/Aeolidiella-papillosa-@-Netherlands-17-03-2017-by-Stefan-Verheyen-2.jpg\" data-orig-size=\"1024,768\" data-comments-opened=\"\" data-image-title=\"Aeolidia papillosa @ Netherlands 17-03-2017 by Stefan Verheyen\" data-image-description=\"&lt;p&gt;Aeolidia papillosa @ Netherlands 17-03-2017 by Stefan Verheyen&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2017\/12\/Aeolidiella-papillosa-%40-Netherlands-17-03-2017-by-Stefan-Verheyen-2.jpg?fit=300%2C225&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2017\/12\/Aeolidiella-papillosa-%40-Netherlands-17-03-2017-by-Stefan-Verheyen-2.jpg?fit=900%2C675&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2017\/12\/Aeolidiella-papillosa-%40-Netherlands-17-03-2017-by-Stefan-Verheyen-2.jpg?w=296&#038;h=222&#038;ssl=1\" width=\"296\" height=\"222\" loading=\"lazy\" data-original-width=\"296\" data-original-height=\"222\" itemprop=\"http:\/\/schema.org\/image\" title=\"Aeolidia papillosa @ Netherlands 17-03-2017 by Stefan Verheyen\" alt=\"Aeolidia papillosa @ Netherlands 17-03-2017 by Stefan Verheyen\" style=\"width: 296px; height: 222px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Aeolidia papillosa @ Netherlands 17-03-2017 by Stefan Verheyen <\/div> <\/div> <\/div> <!-- close group --> <\/div> <!-- close row --> <div class=\"gallery-row\" style=\"width: 900px; height: 451px;\" data-original-width=\"900\" data-original-height=\"451\" > <div class=\"gallery-group images-2\" style=\"width: 300px; height: 451px;\" data-original-width=\"300\" data-original-height=\"451\" > <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/aeolidioidea\/aeolidia-papillosa\/olympus-digital-camera-136\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"296\"> <meta itemprop=\"height\" content=\"222\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"24241\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2017\/12\/Aeolidiella-papillosa-@-Netherlands-17-03-2017-by-Stefan-Verheyen.jpg\" data-orig-size=\"1024,768\" data-comments-opened=\"\" data-image-title=\"Aeolidia papillosa @ Netherlands 17-03-2017 by Stefan Verheyen\" data-image-description=\"&lt;p&gt;Aeolidia papillosa @ Netherlands 17-03-2017 by Stefan Verheyen&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2017\/12\/Aeolidiella-papillosa-%40-Netherlands-17-03-2017-by-Stefan-Verheyen.jpg?fit=300%2C225&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2017\/12\/Aeolidiella-papillosa-%40-Netherlands-17-03-2017-by-Stefan-Verheyen.jpg?fit=900%2C675&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2017\/12\/Aeolidiella-papillosa-%40-Netherlands-17-03-2017-by-Stefan-Verheyen.jpg?w=296&#038;h=222&#038;ssl=1\" width=\"296\" height=\"222\" loading=\"lazy\" data-original-width=\"296\" data-original-height=\"222\" itemprop=\"http:\/\/schema.org\/image\" title=\"Aeolidia papillosa @ Netherlands 17-03-2017 by Stefan Verheyen\" alt=\"Aeolidia papillosa @ Netherlands 17-03-2017 by Stefan Verheyen\" style=\"width: 296px; height: 222px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Aeolidia papillosa @ Netherlands 17-03-2017 by Stefan Verheyen <\/div> <\/div> <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/aeolidioidea\/aeolidia-papillosa\/olympus-digital-camera-137\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"296\"> <meta itemprop=\"height\" content=\"221\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"24242\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2017\/12\/Aeolidiella-papillosa-@-Netherlands-20-02-2017-by-Stefan-Verheyen-2.jpg\" data-orig-size=\"1024,768\" data-comments-opened=\"\" data-image-title=\"Aeolidia papillosa @ Netherlands 20-02-2017 by Stefan Verheyen\" data-image-description=\"&lt;p&gt;Aeolidia papillosa @ Netherlands 20-02-2017 by Stefan Verheyen&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2017\/12\/Aeolidiella-papillosa-%40-Netherlands-20-02-2017-by-Stefan-Verheyen-2.jpg?fit=300%2C225&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2017\/12\/Aeolidiella-papillosa-%40-Netherlands-20-02-2017-by-Stefan-Verheyen-2.jpg?fit=900%2C675&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2017\/12\/Aeolidiella-papillosa-%40-Netherlands-20-02-2017-by-Stefan-Verheyen-2.jpg?w=296&#038;h=221&#038;ssl=1\" width=\"296\" height=\"221\" loading=\"lazy\" data-original-width=\"296\" data-original-height=\"221\" itemprop=\"http:\/\/schema.org\/image\" title=\"Aeolidia papillosa @ Netherlands 20-02-2017 by Stefan Verheyen\" alt=\"Aeolidia papillosa @ Netherlands 20-02-2017 by Stefan Verheyen\" style=\"width: 296px; height: 221px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Aeolidia papillosa @ Netherlands 20-02-2017 by Stefan Verheyen <\/div> <\/div> <\/div> <!-- close group --> <div class=\"gallery-group images-1\" style=\"width: 600px; height: 451px;\" data-original-width=\"600\" data-original-height=\"451\" > <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/aeolidioidea\/aeolidia-papillosa\/olympus-digital-camera-138\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"596\"> <meta itemprop=\"height\" content=\"447\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"24243\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2017\/12\/Aeolidiella-papillosa-@-Netherlands-20-02-2017-by-Stefan-Verheyen.jpg\" data-orig-size=\"1024,768\" data-comments-opened=\"\" data-image-title=\"Aeolidia papillosa @ Netherlands 20-02-2017 by Stefan Verheyen\" data-image-description=\"&lt;p&gt;Aeolidia papillosa @ Netherlands 20-02-2017 by Stefan Verheyen&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2017\/12\/Aeolidiella-papillosa-%40-Netherlands-20-02-2017-by-Stefan-Verheyen.jpg?fit=300%2C225&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2017\/12\/Aeolidiella-papillosa-%40-Netherlands-20-02-2017-by-Stefan-Verheyen.jpg?fit=900%2C675&#038;ssl=1\" src=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2017\/12\/Aeolidiella-papillosa-%40-Netherlands-20-02-2017-by-Stefan-Verheyen.jpg?w=596&#038;h=447&#038;ssl=1\" width=\"596\" height=\"447\" loading=\"lazy\" data-original-width=\"596\" data-original-height=\"447\" itemprop=\"http:\/\/schema.org\/image\" title=\"Aeolidia papillosa @ Netherlands 20-02-2017 by Stefan Verheyen\" alt=\"Aeolidia papillosa @ Netherlands 20-02-2017 by Stefan Verheyen\" style=\"width: 596px; height: 447px;\" 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loading=\"lazy\" data-original-width=\"446\" data-original-height=\"335\" itemprop=\"http:\/\/schema.org\/image\" title=\"Aeolidia papillosa @ Netherlands 26-01-2017 by Stefan Verheyen\" alt=\"Aeolidia papillosa @ Netherlands 26-01-2017 by Stefan Verheyen\" style=\"width: 446px; height: 335px;\" \/> <\/a> <div class=\"tiled-gallery-caption\" itemprop=\"caption description\"> Aeolidia papillosa @ Netherlands 26-01-2017 by Stefan Verheyen <\/div> <\/div> <\/div> <!-- close group --> <div class=\"gallery-group images-1\" style=\"width: 450px; height: 339px;\" data-original-width=\"450\" data-original-height=\"339\" > <div class=\"tiled-gallery-item tiled-gallery-item-large\" itemprop=\"associatedMedia\" itemscope itemtype=\"http:\/\/schema.org\/ImageObject\"> <a href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/aeolidioidea\/aeolidia-papillosa\/olympus-digital-camera-235\/\" border=\"0\" itemprop=\"url\"> <meta itemprop=\"width\" content=\"446\"> <meta itemprop=\"height\" content=\"335\"> <img decoding=\"async\" class=\"\" data-attachment-id=\"27973\" data-orig-file=\"https:\/\/opistobranquis.info\/wp-content\/uploads\/2019\/01\/Aeolidia-papillosa-@-Netherlands-29-03-2018-by-Stefan-Verheyen.jpg\" data-orig-size=\"2114,1586\" data-comments-opened=\"\" data-image-title=\"Aeolidia papillosa @ Netherlands 29-03-2018 by Stefan Verheyen\" data-image-description=\"&lt;p&gt;Aeolidia papillosa @ Netherlands 29-03-2018 by Stefan Verheyen&lt;\/p&gt; \" data-medium-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2019\/01\/Aeolidia-papillosa-%40-Netherlands-29-03-2018-by-Stefan-Verheyen.jpg?fit=300%2C225&#038;ssl=1\" data-large-file=\"https:\/\/i0.wp.com\/opistobranquis.info\/wp-content\/uploads\/2019\/01\/Aeolidia-papillosa-%40-Netherlands-29-03-2018-by-Stefan-Verheyen.jpg?fit=900%2C675&#038;ssl=1\" 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Verhandlungen der koniglich-kaiserlich Zoologisch-botanischen Gesellschaft in Wien (Abhandlungen) 24:395-416, pls. 8-11.<\/div>\n<\/div>\n\t\t\t\t<\/div><!-- .zp-Entry .zpSearchResultsItem -->\t\t\t\t<div id=\"zp-ID-8355-1246567-V2K73CV7\" data-zp-author-date='Bergh-1894' data-zp-date-author='1894-Bergh' data-zp-date='1894' data-zp-year='1894' data-zp-itemtype='journalArticle' class=\"zp-Entry zpSearchResultsItem\">\n<div class=\"csl-bib-body\" style=\"line-height: 1.35; padding-left: 1em; text-indent:-1em;\">\n  <div class=\"csl-entry\">Bergh, L. S. R. (1894). Die Opisthobranchien. Reports on the dredging operations off the west coast of Central America to the Galapagos, to the west coast of Mexico, and in the Gulf of California, in charge of Alexander Agassiz, carried on by the U.S. Fish Commission steamer \u201cAlbatross,\u201d during 1891, Lieut. Commander Z. L. Tanner, U.S.N. commanding. Part 13. <i>Bulletin of the Museum of Comparative Zoology 25: 125\u2013233. [October 1894]<\/i>.<\/div>\n<\/div>\n\t\t\t\t<\/div><!-- .zp-Entry .zpSearchResultsItem -->\n\t\t\t<\/div><!-- .zp-zp-SEO-Content -->\n\t\t<\/div><!-- .zp-List -->\n\t<\/div><!--.zp-Zotpress-->\n\n<p><\/p>Bibliography based on the works by <b>Steve Long, 2006<\/b>. <a target=_blank href=https:\/\/opistobranquis.info\/wp-content\/uploads\/2016\/06\/Long-S.-2006-Bibliography-of-Opisthobranchia-1554-2000.pdf>Bibliography of Opisthobranchia 1554-2000<\/a> and <b>Gary McDonald, 2009<\/b>. <a href=http:\/\/escholarship.org\/uc\/item\/8115h0wz>Bibliographia Nudibranchia<\/a>, with later updates from other resources.<p><\/p><\/ul><p><\/p><p><strong>Further reading<\/strong><\/p><span style=\"font-size:10pt\"><ul><ul class=\"blogroll-links\"><li><a href=\"http:\/\/www.asturnatura.com\/especie\/aeolidia-papillosa.html\">AsturNatura<\/a><\/li><li><a href=\"http:\/\/www.biodiversitylibrary.org\/name\/Aeolidia%20papillosa\">Biodiversity Heritage Library<\/a><\/li><li><a href=\"http:\/\/doris.ffessm.fr\/fiche2.asp?fiche_numero=150\">DORIS<\/a><\/li><li><a href=\"http:\/\/nature22.com\/estran22\/mollusques\/gasteropodesbis\/gasteropodes5.html\">Estran 22 - Faune et flore de la zone de balancement des mar\u00e9es en C\u00f4tes d&#039;Armor<\/a><\/li><li><a href=\"https:\/\/www.flickr.com\/photos\/56388191@N08\/sets\/72177720302685814\/\">European Sea Slugs by Morddyn<\/a><\/li><li><a href=\"https:\/\/www.flickr.com\/search\/?text=Aeolidia%20papillosa\">Flickr pictures<\/a><\/li><li><a href=\"http:\/\/www.irmng.org\/aphia.php?p=taxlist&#038;tName=Aeolidia%20papillosa\">Interim Register of Marine and Nonmarine Genera<\/a><\/li><li><a href=\"http:\/\/www.ncbi.nlm.nih.gov\/Taxonomy\/Browser\/wwwtax.cgi?mode=Undef&#038;name=Aeolidia%20papillosa&#038;lvl=0&#038;srchmode=1\">NCBI GenBank<\/a><\/li><li><a href=\"http:\/\/iobis.org\/mapper\/?taxon=Aeolidia%20papillosa\">OBIS - Search by Taxon<\/a><\/li><li><a href=\"http:\/\/www.nudibranch.org\/Scottish%20Nudibranchs\/aeolidia-papillosa.html\">Scottish Nudibranchs<\/a><\/li><li><a href=\"http:\/\/www.seaslugforum.net\/showall\/aeolpapi\">Sea Slug Forum<\/a><\/li><li><a href=\"http:\/\/www.conchsoc.net\/spaccount\/Aeolidia-papillosa\">The Conchological Society of Great Britain and Ireland<\/a><\/li><li><a href=\"http:\/\/slugsite.us\/bow\/nudwk407.htm\">The Slug Site<\/a><\/li><li><a href=\"http:\/\/www.marinespecies.org\/aphia.php?p=taxdetails&#038;id=138709\">World Register of Marine Species<\/a><\/li><\/ul><\/ul><\/span><p><strong>Cite this article as: <\/strong><\/p><div style=\"display: block; background: #444444; padding: 10px 30px 10px 10px; border-radius: 5px; line-height: 18px;\"><p class=\"citacio\"><b>Pontes, Miquel, Manuel Ballesteros, Enric Madrenas<\/b> (2025) <em>\"Aeolidia papillosa\"<\/em> in <em>OPK-Opistobranquis<\/em>. Published: 22\/09\/2013. Accessed: 27\/07\/2026. Available at (<a href=https:\/\/opistobranquis.info\/en\/?p=8355>https:\/\/opistobranquis.info\/en\/?p=8355<\/a>)<\/p><p><\/p><\/div><p><\/p><\/p>","protected":false},"excerpt":{"rendered":"<p>Aeolidia papillosa (Linnaeus, 1761) Synonyms Aeolidia herculea Troschel, 1866 Aeolidia papillosa var. pacifica Bergh, 1879 Aeolidia serotina Bergh, 1873 Aeolis lesliana MacGillivray, 1843 Aeolis murrayana MacGillivray, 1843 Doris bodoensis Gunnerus, 1770 Doris vermigera Turton, 1807 Eolidia zetlandica Forbes &amp; Goodsir, 1839 Eolis campbellii Cunningham, 1871 Eolis cuvieri Lamarck, 1819 Eolis farinacea Stimpson, 1853 Eolis obtusalis Alder &amp; Hancock, 1842 Eolis papillosa (Linnaeus, 1761) Eolis papillosa var. albina Dautzenberg &amp; Durouchoux,\u2026<\/p>\n<p class=\"continue-reading-button\"> <a class=\"continue-reading-link\" href=\"https:\/\/opistobranquis.info\/en\/guia\/nudibranchia\/aeolidioidea\/aeolidia-papillosa\/\">Continue reading<i class=\"crycon-right-dir\"><\/i><\/a><\/p>\n","protected":false},"author":423,"featured_media":11237,"parent":456,"menu_order":8137,"comment_status":"closed","ping_status":"closed","template":"","meta":{"footnotes":""},"class_list":["post-8355","page","type-page","status-publish","has-post-thumbnail","hentry"],"jetpack_sharing_enabled":true,"_links":{"self":[{"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages\/8355","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/users\/423"}],"replies":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/comments?post=8355"}],"version-history":[{"count":1,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages\/8355\/revisions"}],"predecessor-version":[{"id":25536,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages\/8355\/revisions\/25536"}],"up":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/pages\/456"}],"wp:featuredmedia":[{"embeddable":true,"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/media\/11237"}],"wp:attachment":[{"href":"https:\/\/opistobranquis.info\/en\/wp-json\/wp\/v2\/media?parent=8355"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}