Microchlamylla gracilis

Microchlamylla gracilis  (Alder & Hancock, 1844)

Nota taxonómica: La clasificación de los Flabellinidae había permanecido bastante estable hasta que en 2017 aparecieron una serie de trabajos (Furfaro et al. 2017; Korshunova et al. 2017) que pretendían aclarar el estatus de la familia Flabellinidae. El objetivo principal del trabajo de Furfaro et al. era caracterizar molecularmente las especies mediterráneas mientras que el trabajo de Korshunova et al. pretendía ahondar en las relaciones filogenéticas entre diversos miembros de la familia Flabellinidae y las demás familias de los aeolidáceos.
Ambos trabajos se basaban en la combinación de técnicas moleculares y morfologicas y, de hecho, no ofrecen resultados muy distintos, pero sí difieren en el tamaño y procedencia de las muestras estudiadas y, sobre todo, en la interpretación de dichos resultados. Tras la aparición del trabajo de Furfaro et al., muchas especies Mediterráneas de los géneros Calmella, Flabellina y Piseinotecus se agruparon bajo el género común Flabellina, pero tuvieron determinados problemas con algunas especies que no encajaban bien con la clasificación propuesta (p.e. Flabellina babai), descubrieron que las poblaciones mediterránea y atlántica de Flabellina ischitana corresponden en realidad a dos especies cripticas diferentes, y constataron los problemas del grupo críptico Calmella cavolini / Flabellina confusa / Piseinotecus gaditanuspor lo que indicaron la necesidad de nuevos estudios para aclarar su estatus.
Curiosamente estos estudios se estaban llevando prácticamente en paralelo por el grupo de Korshunova et al. pero sobre una muestra de especies mucho más amplia y que comprendía ejemplares del Ártico, Atlántico norte, océano Pacífico e Índico. Este segundo trabajo confirma la polifília de la familia Flabellinidae, pero su forma de interpretar los resultados representa una verdadera revolución para la taxonomía de los aeolidáceos, especialmente para la familia Flabellinidae.
Ambos trabajos observan que hay dos clados (grupos) bien diferenciados de Flabellinidae: el de Coryphella pedata y similares, con cerata que salen directamente del dorso, y el de Flabellina affinis y similares, que tienen los cerata de cada grupo pedunculados. Aunque Furfaro et al. consideran a las especies de ambos clados pertenecientes al género Flabellina dentro de la familia Flabellinidae, Korshunova et al. diferencia dos familias: Coryphellidae y Flabellinidae sensu stricto, creando además multitud de géneros diferentes en estas familias para incluir a las especies que estudian. Su propuesta taxonómica, curiosamente, da respuesta a los problemas hallados por Furfaro et al.
En cierta manera ambos trabajos se complementan, si bien en el trabajo de Korshunova et al. se evidencia la falta de estudios sobre las especies de flabellinidos tropicales y del sur de América y África, por lo que aún no se ha dicho la última palabra. La propuesta de creación de nuevos géneros para reunir pequeños grupos de especies, en vez de géneros multiespecíficos, parece ser la tendencia en algunos de los trabajos filogenéticos de los últimos años. Puede consultarse una discusión detallada de esta apasionante controversia en (https://opistobranquis.info/es/flabellinidae/).

• Coryphella gracilis (Alder & Hancock, 1844)
• Coryphella rufibranchialis var. clavigera Odhner, 1929
• Eolis gracilis Alder & Hancock, 1844 (original)
• Eolis smaragdina Alder & Hancock, 1851
• Flabellina gracilis (Alder & Hancock, 1844)

Descripción

Biología

Etimología

• Microchlamylla. Derivado de “micro” y “chlamylla“. En referencia a los conductos deferentes (vas deferens) inusualmente largos de este género, típicos del género Chlamylla de la familia Paracoryphellidae, pero con un menor tamaño corporal (“micro“) y con el borde del notum discontínuo, común a la família Coryphellidae.
• Gracilis. Del Latín “gracilis”, delgado, menudo.

Distribución

Citas georeferenciadas conocidas de la especie: Microchlamylla gracilis

Fonts:
OBIS GROC 2010-2011 Enric Madrenas João Pedro Silva Bernard Picton GBIF	OPK VIMAR Manuel Ballesteros M@re Nostrum Altres fonts Marine Regions

Otras fotos

Microchlamylla gracilis (Sweden) by Klas Malmberg

Microchlamylla gracilis @ Netherlands 17-02-2017 by Stefan Verheyen

Microchlamylla gracilis @ Netherlands by Stefan Verheyen

Microchlamylla gracilis @ Netherlands by Stefan Verheyen

Microchlamylla gracilis @ Netherlands 17-02-2017 by Stefan Verheyen

Microchlamylla gracilis @ Netherlands 24-03-2017 by Stefan Verheyen

Microchlamylla gracilis feeding on Eudendrium @ Netherlands 21-04-2017 by Stefan Verheyen

Microchlamylla gracilis with spawn @ Netherlands 27-03-2017 by Stefan Verheyen

Microchlamylla gracilis @ Netherlands 29-03-2018 by Stefan Verheyen

1246567 Microchlamylla gracilis 1 eunomia-revista-en-cultura-de-la-legalidad 50 creator asc 1 16020 https://opistobranquis.info/wp-content/plugins/zotpress/
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Bibliografía basada en los trabajos de Steve Long, 2006. Bibliography of Opisthobranchia 1554-2000 y Gary McDonald, 2009. Bibliographia Nudibranchia, con actualizaciones posteriores procedentes de otras fuentes.

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