Los Cylichnoidea son una superfamilia de moluscos gasterópodos heterobranquios marinos que incluye especies generalmente pequeñas, caracterizadas a menudo por una concha cilíndrica o alargada, con una espira muy baja o casi invisible. Su forma compacta y la reducción de la espira les confieren un aspecto particular, que permite distinguir a muchos de sus representantes de otros gasterópodos con conchas más globosas o cónicas. Aunque a menudo pasan desapercibidos, constituyen un componente interesante de la diversidad de los pequeños moluscos que habitan los fondos marinos.
Los Cylichnoidea viven principalmente en ambientes bentónicos, desde las aguas litorales poco profundas hasta zonas de mayor profundidad. Se pueden encontrar en sedimentos arenosos o fangosos, donde muchas especies se entierran parcialmente o se desplazan entre los granos del sustrato. Este estilo de vida, junto con sus reducidas dimensiones, hace que sean difíciles de detectar durante las observaciones submarinas convencionales. En cambio, pueden aparecer en muestras de sedimento obtenidas mediante técnicas de muestreo bentónico.
La concha es uno de los elementos más útiles para reconocer este grupo. En muchas especies es alargada, cilíndrica o ligeramente ovalada, con una abertura estrecha y prolongada que puede ocupar buena parte de la longitud de la concha. La espira puede estar muy reducida o hundida, mientras que la superficie externa puede ser lisa o presentar estrías y otras ornamentaciones finas. A pesar de esta apariencia relativamente uniforme, el estudio detallado de la forma, las proporciones y los caracteres anatómicos permite diferenciar especies que, a primera vista, pueden parecer muy similares.
En cuanto a la alimentación, numerosos representantes de este grupo están asociados a los sedimentos y aprovechan los recursos disponibles en estos ambientes. Algunas especies se alimentan de pequeños organismos o de material orgánico presente en el sustrato, aunque los hábitos tróficos pueden variar entre los distintos linajes. De este modo, participan en las complejas redes alimentarias de las comunidades bentónicas, especialmente en los ecosistemas dominados por sedimentos blandos.
La clasificación de los Cylichnoidea y las relaciones evolutivas entre sus representantes han sido objeto de revisión a medida que se han incorporado datos anatómicos y moleculares. La semejanza externa entre muchas especies no siempre refleja una relación evolutiva estrecha, lo que pone de manifiesto la importancia de combinar diferentes tipos de caracteres para comprender la diversificación de los heterobranquios.
A pesar de su diminuto tamaño y sus hábitos discretos, los Cylichnoidea son un grupo relevante para estudiar la biodiversidad de los sedimentos marinos. Su morfología característica, su adaptación a la vida bentónica y las dificultades que plantea su identificación los convierten en un ejemplo interesante de la diversidad menos visible de los gasterópodos marinos.
La taxonomía actual según WoRMS es:
Especies de Cylichnoidea citadas en el Mediterráneo o en la Península Ibérica:
Cylichna alba
Cylichna brocchii
Cylichna cylindracea
Diaphana globosa
Diaphana minuta
Mnestia girardi
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Bibliografía basada en los trabajos de Steve Long, 2006. Bibliography of Opisthobranchia 1554-2000 y Gary McDonald, 2009. Bibliographia Nudibranchia, con actualizaciones posteriores procedentes de otras fuentes.
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