Acochlidiimorpha

Se trata de un pequeño grupo de Heterobranquios que comprende unas 30 especies en todo el mundo y que viven en el medio intersticial marino o de agua dulce. Están muy poco estudiados, son de muy pequeño tamaño y tienen una morfología corporal algo aberrante para tratarse de moluscos y que está adaptada para vivir y desplazarse entre las partículas de sedimento.

Acochlidium fijiiensis Haynes & Kenchington, 1991
http://www.wikipedia.org

Tienen un cuerpo alargado, sin concha en estado adulto y un manto que suele disponer de numerosas espículas. No tienen ni cavidad paleal ni branquias y la cabeza dispone de tentáculos orales y rinóforos. Generalmente los sexos son separados. Posiblemente se alimentan de microorganismos que viven en el medio intersticial.

Considerados tradicionalmente como un grupo peculiar de Opistobranquios, los trabajos recientes de Jorger et al. (2010) y de Schrödl et al. (2011) tienden a acercarlos a los gasterópodos Pulmonados, dentro del clado de los Panpulmonata.

La taxonomía de los Acochlidiimorpha según el trabajo de Bouchet et al. (2017) es la siguiente:

• Superorden Acochlidiimorpha
  • Superfamilia Acochlidioidea Küthe, 1935
    • Familia Acochlidiidae Küthe, 1935
      • Género Acochlidium Strubell, 1892
      • Género Palliohedyla Rankin, 1979
      • Género Palliohedyle Rankin, 1979
      • Género Strubellia Odhner, 1937
    • Familia Aitengidae Swennen & Buatip, 2009
      • Género Aiteng Swennen & Buatip, 2009
    • Familia Bathyhedylidae Neusser, Jörger, Lodde-Bensch, Strong & Schrödl, 2016
      • Género Bathyhedyle Neusser, Jörger, Lodde-Bensch, Strong & Schrödl, 2016
    • Familia Hedylopsidae Odhner, 1952
      • Género Hedylopsis Thiele, 1931
    • Familia Pseudunelidae Rankin, 1979
      • Género Pseudunela Salvini-Plawen, 1973
    • Familia Tantulidae Rankin, 1979
      • Género Tantulum Rankin, 1979
  • Superfamilia Parhedyloidea Thiele, 1931
    • Familia Asperspinidae Rankin, 1979
      • Género Asperspina Rankin, 1979
      • Género Minicheviella Starobogatov, 1983
    • Familia Parhedylidae Thiele, 1931
      • Género Microhedyle Hertling, 1930
      • Género Parhedyle Thiele, 1931
      • Género Pontohedyle Golikov & Starobogatov, 1972

Pontohedyle milaschewitchii (Kowalevsky, 1901)
A) Esquema de un ejemplar entero en vista dorsal (dg: glándula digestiva; ey: ojos;  lt: tentáculo labial; sp1 y sp2: espículas; vs: saco visceral)
B, C) Fotografias ejemplar vivo mostrando la variación de su forma externa en movimiento. (Jorger et al, 2008)

En el Mediterráneo y en la Península Ibérica se han citado estas especies:

Asperspina loricata	Hedylopsis spiculifera
Microhedyle glandulifera	Pontohedyle milaschewitchii

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Bibliografía basada en los trabajos de Steve Long, 2006. Bibliography of Opisthobranchia 1554-2000 y Gary McDonald, 2009. Bibliographia Nudibranchia, con actualizaciones posteriores procedentes de otras fuentes.