Piseinotecus soussi

Piseinotecus soussi Tamsouri, Carmona, Moukrim & Cervera, 2014

Piseinotecus soussi by Enric Madrenas










































Piseinotecus soussi  Tamsouri, Carmona, Moukrim & Cervera, 2014

 Classification according to Bouchet et al. (2017)
Taxonomic source: World Register of Marine Species (AphiaID: 818876).
Adult animals of this recently described species measure between 8 and 13 mm in length. The body is narrow and has a long, pointed tail when the animal is moving. The general body color is pink or bluish-pink. The oral tentacles and rhinophores have approximately the same size and are of the same color of the body, but with whitish tips because on many white granulations on this part. The rhinophores are smooth. The eyes are barely visible in the rear base of the rhinophores. Depending on the size of the specimens there are 7 to 9 groups of cerata where each may have from 13 cerata in the first group to only 1 in the last one. The cerata from each group come from a common base or stalk, as in some species of Flabellina. The cerata are cylindrical, relatively long and with a pointed tip; they are semitransparent and have the same coloration of the body but the more or less dark brown digestive gland can be observed inside, filling most of the cerata. The subapical area of the cerata usually have small white spots or scores and the tip is whitish, where the cnidosac can be observed. The foot is narrow and two short, triangular propodials tentacles can be observed in its anterior side. The gonopore is located right under the first group of cerata while the anus is dorsal (acleioproctal position).

So far this species has been found living under rocks in the intertidal ponds or in the subtidal stratum in rock walls with an abundance of algae and hydroids, on which it may feed. Nothing else is known about his spawn nor other details of its biology.


  • Piseinotecus means “I stepped on Teco.” Teco was a dog belonging to Ernst and Eveline Marcus. Apparently Prof. Diva Diniz Corrêa, friend of the family, unadvertedly stepped on the dog and while the Marcuses were looking for a new generic name she said in Portuguese: “Pisei”=I stteped, “no”=onto, “Tecus”=the dog’s name. Later on that name became also a family name: Piseinotecidae (ref:Curious Taxonomy and Marcus, Ev. 1987).
  • Soussi. Related to Souss Massa Drâa (in Agadir province), where most specimens of this new species have been collected.

This species is known from the Atlantic coast of Morocco (type locality of this species is Cap Ghir, about 25 km from Agadir), the coast of Cadiz, southern Italy, Banyuls-sur-Mer (France) and the coast of Catalonia, where it has been observed in Caials (Cadaqués), Cala Jóncols (Roses), the Medes Islands and Tossa de Mar. It may have a wider distribution range because it has been confused with species of the genus Flabellina (F. pedata, F. affinis, F. ischitana) because of their similar morphology and coloration.

Known georeferenced records of the species: Piseinotecus soussi
: GROC 2010-2011
: Enric Madrenas
: João Pedro Silva
: Bernard Picton
: Manuel Ballesteros.
: M@re Nostrum
: Altres fonts
: Marine Regions

Similar species
Edmundsella pedata almost identical but without white spots on cerata.


    Western Mediterranean:1 out of 5 stars
    Eastern Mediterranean:0 out of 5 stars
    Atlantic Ocean:1 out of 5 stars

This chart displays the monthly observation probability for Piseinotecus soussi based on our own records.

More pictures


    Ballesteros, M., E. Madrenas, and M. Pontes. 2021. OPK - Opistobranquis. (https://opistobranquis.info/).
    Ballesteros, M., E. Madrenas, and M. Pontes. 2016. Actualización del catálogo de los moluscos opistobranquios (Gastropoda: Heterobranchia) de las costas catalanas. Spira 6(2016): 1-28.
    Ballesteros, M., M. Pontes, and E. Madrenas. 2015. Opisthobranch from Medes Islands (Marine Protected Area, Costa Brava, Catalonia, NE Spain): 40 years of study.
    Ballesteros, M., M. Pontes, and E. Madrenas. 2019. Els nudibranquis del mar català. Brau, Figueres.
    Flanders Marine Institute. 2018. Maritime Boundaries Geodatabase: Territorial Seas. Maritime Boundaries Geodatabase. (http://www.marineregions.org/).
    Furfaro, G., and P. Mariottini. 2019. Less rare than we thought: two new localities for Piseinotecus soussi Tamsouri, Carmona, Moukrim & Cervera, 2014 along the Tyrrhenian coast. Tübitak - Turkish Journal of Zoology 43: 287-289.
    Furfaro, G., D. Salvi, E. Mancini, and P. Mariottini. 2017. A multilocus view on Mediterranean aeolid nudibranchs (Mollusca): Systematics and cryptic diversity of Flabellinidae and Piseinotecidae. Molecular Phylogenetics and Evolution 118: 13–22.
    Furfaro, G., F. Vitale, C. Licchelli, and P. Mariottini. 2020. Two Seas for One Great Diversity: Checklist of the Marine Heterobranchia (Mollusca; Gastropoda) from the Salento Peninsula (South-East Italy). Diversity. 12: 171.
    Long, S. J. 2006. Bibliography of Opisthobranchia 1554-2000. Bayside Books & Press, Tustin, CA, U.S.A. 672p.
    Marcus, Ev. d. B. R. 1987. Selected recollections from my life. American Malacological Bulletin 5(2):183-184.
    Martínez-Chacón, M. 2018. Nudibranquios de la isla de Tarifa, pequeñas joyas del estrecho de Gibraltar. Junta de Andalucía. Consejería de Medio Ambiente y Ordenación del Territorio.
    McDonald, G. 2009. Bibliographia Nudibranchia. 2nd Online Edition, Annotated. 1072 pp  Institute of Marine Sciences, University of California, Santa Cruz. (http://escholarship.org/uc/item/8115h0wz).
    Prkić, J., A. Petani, Ð. Iglić, and L. Lanča. 2018. Stražnjoškržnjaci Jadranskoga Mora: Slikovni Atlas i Popis Hrvatskih Vrsta / Opisthobranchs of the Adriatic Sea: Photographic Atlas and List of Croatian Species. Ronilaćki Klub Sveti Roko, Bibinje.
    Tamsouri, N., L. Carmona, A. Moukrim, and J. L. Cervera. 2014. Description of a new species of Piseinotecus (Gastropoda, Heterobranchia, Piseinotecidae) from the northeastern Atlantic Ocean. Bulletin of Marine Science. 90(4): 991–997.
    Tamsouri, N., L. Carmona, A. Moukrim, and J. L. Cervera. 2014. Addenda to the article Bull Mar Sci. 90(4):991–997, 2014: Description of a new species of Piseinotecus (Castropoda, Heterobranchia, Piseinotecidae) from the northeastern Atlantic Ocean. Bulletin of Marine Science. 91(1): 83–84.
    WoRMS Editorial Board. 2021. World Register of Marine Species. WoRMS. (http://www.marinespecies.org).

    Bibliography based on the works by Steve Long, 2006. Bibliography of Opisthobranchia 1554-2000 and Gary McDonald, 2009. Bibliographia Nudibranchia, with later updates from other resources.

Further reading

Cite this article as:

Cervera, Lucas, Manuel Ballesteros, Enric Madrenas, Miquel Pontes (2021) "Piseinotecus soussi" in OPK-Opistobranquis. Published: 02/10/2014. Accessed: 24/01/2021. Available at (https://opistobranquis.info/en/lbpXU)

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