Doto verdicioi

Doto verdicioi  Ortea & Urgorri, 1978

Doto verdicioi @ Lapamán (Ría de Pontevedra, Spain) 28.03.1979 by Victoriano Urgorri

Class: Gastropoda  Cuvier, 1797
Subclass: Heterobranchia  J.E. Gray, 1840
Clade: Euthyneura  Spengel, 1881
Superorder: Nudipleura  Wägele & Willan, 2000
Order: Nudibranchia  Cuvier, 1817
Suborder: Dexiarchia  Schrödl, Wägele & Willan, 2001
Infraorder: Cladobranchia Willan and Morton, 1984
Parvorder: [unassigned] Cladobranchia Willan & Morton, 1984
Family: Dotidae  J.E. Gray, 1853
Genus: Doto  Oken, 1815
Species: Doto verdicioi  Ortea & Urgorri, 1978

Body color usually semi-transparent white, pigmented with dark red spots on the back and flanks, leaving depigmented areas between the cerata. The reddish pigmentation forms a very tight band that runs along the back from the head to the last cerata, becoming disperse on the tail. Dotting on the sides is very thin and scattered. Gonad is visible by transparency as a pink cream organ and is located between the first and fifth cera. The rhinophores are approximately 0.9mm long, pigmented with large red spots, with a white stained apex. The rhinophoric sheath measures 0.5mm and has some white scores on the outside, sometimes with one or two red spots, but inside has large red spots. Cephalic lobes are inconspicuous and only sometimes have some reddish dots. The base of the cerata and the areas between them are depigmented. The dorsum has five or six cerata on each side, each cera with two rows of four slightly prominent blunt tubercles. These tubercles are white inside with a white apex without dark spots, but have numerous red spots surrounding the apex and irregularly distributed along the rest of the cera. The apical tubercle is much larger than the others, somewhat chubby, with a hemispherical apex with internal white spots; subterminally it presents a more or less defined ring formed by independent reddish spots that in the larger specimens becomes almost continuous. The interior of the cerata is pink with small white “horseshoes”. The pseudobranch is very characteristic when it is well developed and consists of five processes, one apical and two on the sides, the ones on the sides very large and flattened so they protrude on the sides of the cera. The anal papilla is prominent, dotted in white and it is located between the first two cerata on the right side of the body. The genital orifice is located somewhat ahead of the first cera on the right side of the body, generally in a highly pigmented red zone.

The specimens have been found on hidrarians of the genus Aglaophenia, in walls with relatively high hydrodynamics. Spawning ribbon forms two or three corrugations, although single horseshoe shaped ribbons are also common.


  • Doto, from Latin “Doto”, a sea nymph, from Greek “Dōtō”
  • Verdicioi, dedicated to the village where this species was first collected, Verdicio (Peñas cape, Asturias, Spain)

Species basically cited on the Atlantic coast of the Iberian Peninsula, in Asturias, Galicia and Portugal.

Known georeferenced records of the species: Doto verdicioi
: GROC 2010-2011
: Enric Madrenas
: João Pedro Silva
: Bernard Picton
: Manuel Ballesteros.
: M@re Nostrum
: Altres fonts
: Marine Regions


    Western Mediterranean:
    Eastern Mediterranean:
    Atlantic Ocean:

More pictures


Further reading

Cite this article as:

Ballesteros, Manuel, Enric Madrenas, Miquel Pontes (2012-2018) "Doto verdicioi" in OPK-Opistobranquis, Published: 24/04/2016, Accessed: 17/12/2018 at (

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